# Cornuta

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{{Short description|Extinct order of marine invertebrates}}
{{About|the prehistoric echinoderm taxon|the prehistoric fish taxon|Cornuata}}
{{Automatic taxobox
| fossil_range = [Middle&nbsp;Cambrian](/source/Middle%26nbsp%3BCambrian)–[Late&nbsp;Ordovician](/source/Late%26nbsp%3BOrdovician) ([Rawtheyan](/source/Katian))<ref name=PBDB>{{Cite PBDB|taxon|31154|Cornuta}}</ref> {{Geological range|Middle Cambrian|Katian|}}
| image = Cothurnocystis.jpg
| image_caption = Reconstruction of the cornute ''[Cothurnocystis](/source/Cothurnocystis)''
| image2 = Cothurnocystis elizae CRF.jpg
| image2_caption = Fossil of ''[Cothurnocystis](/source/Cothurnocystis) elizae''
| taxon = Cornuta
| authority = Jaekel, 1901
| subdivision_ranks = Taxa
| subdivision = See Taxa
}}

'''Cornuta''' is an extinct order of [echinoderm](/source/echinoderm)s.  Along with the [mitrate](/source/mitrate)s, they form the [Stylophora](/source/Stylophora).

Their first (probable) representative is ''[Ponticulocarpus](/source/Ponticulocarpus)'' from the Spence Shale (mid Cambrian);<ref>{{cite journal | url = https://www.cambridge.org/core/journals/journal-of-paleontology/article/abs/ponticulocarpus-a-new-cornutegrade-stylophoran-from-the-middle-cambrian-spence-shale-of-utah/FA9710BCB74ED37D3C91B9FAA39BB86D| doi = 10.1017/S0022336000040725| title = ''Ponticulocarpus,'' a new cornute-grade stylophoran from the Middle Cambrian Spence Shale of Utah| year = 1999| last1 = Sumrall| first1 = Colin D.| last2 = Sprinkle| first2 = James| journal = Journal of Paleontology| volume = 73| issue = 5| pages = 886–891| bibcode = 1999JPal...73..886S| s2cid = 132521457| url-access = subscription}}</ref> Ordovician examples also exist.<ref name=PBDB/>

== Taxa ==
As of 2002, the following taxa were recognized as being included in the order Cornuta:<ref name=Dominguez_et_al2002>{{Cite journal |last=Domínguez Alonso |first=Patricio |last2=Jefferies |first2=Richard P. S. |last3=Gil Cid |first3=María Dolores |date=December 2002 |title=An annotated check-list of genera and species of carpoids |url=https://hdl.handle.net/20.500.14352/56880 |journal=Coloquios de Palcontologia |volume=53 |pages=33-68 |issn=1132-1660}}</ref>

{{div col|colwidth=25em|small=yes}}
*''[Acuticarpus](/source/Acuticarpus)'' {{Small|Sumrall, Sprinkle & Guensburg, 1997}}
*''[Amygdalotheca](/source/Amygdalotheca)'' {{Small|Ubaghs, 1969}}
*''[Ampelocarpus](/source/Ampelocarpus)'' {{Small|Lefebvre & Vizcaïno, 1999}}
*''[Arauricystis](/source/Arauricystis)'' {{Small|Lefebvre & Vizcaïno, 1999}}
*''[Archaeocothurnus](/source/Archaeocothurnus)'' {{Small|Sumrall, Sprinkle & Guensburg, 1997}}
*''[Babinocystis](/source/Babinocystis)'' {{Small|Rozhnov, 1990}}
*''[Beryllia](/source/Beryllia_(cornute))'' {{Small|Cripps & Daley, 1994}}
*''[Bohemiaecystis](/source/Bohemiaecystis)'' {{Small|Caster, 1967}}
*''[Ceratocystis](/source/Ceratocystis_(cornute))'' {{Small|Jaekel, 1901}}
*''[Chauvelicystis](/source/Chauvelicystis)'' {{Small|Ubaghs, 1969}}
*''[Cothurnocystis](/source/Cothurnocystis)'' {{Small|Bather, 1913}}
*''[Domfrontia](/source/Domfrontia)'' {{Small|Cripps & Daley, 1994}}
*''[Drepanocarpos](/source/Drepanocarpos)'' {{Small|Smith & Jell, 1999}}
*''[Flabellicystis](/source/Flabellicystis)'' {{Small|Martí Mus, 2002}}
*''[Galliaecystis](/source/Galliaecystis)'' {{Small|Ubaghs, 1969}}
*''[Hanusia](/source/Hanusia_(cornute))'' {{Small|Cripps, 1989}}
*''[Juliaecarpus](/source/Juliaecarpus)'' {{Small|Ruta, 1999}}
*''[Lobocarpus](/source/Lobocarpus)'' {{Small|Ubaghs, 1998}}
*''[Lyricocarpus](/source/Lyricocarpus)'' {{Small|Ubaghs, 1994}}
*''[Milonicystis](/source/Milonicystis)'' {{Small|Chauvel, 1986}}
*''[Nanocarpus](/source/Nanocarpus)'' {{Small|Ubaghs, 1991}}
*''[Nevadaecystis](/source/Nevadaecystis)'' {{Small|Ubaghs, 1967}}
*''[Phyllocystis](/source/Phyllocystis)'' {{Small|Thoral, 1935}}
*''[Ponticulocarpus](/source/Ponticulocarpus)'' {{Small|Sumrall & Sprinkle, 1999}}
*''[Prochauvelicystis](/source/Prochauvelicystis)'' {{Small|Daley, 1992}}
*''[Procothurnocystis](/source/Procothurnocystis)'' {{Small|Woods & Jefferies, 1992}}
*''[Progalliacystis](/source/Progalliacystis)'' {{Small|Cripps, 1989}}
*''[Prokopicystis](/source/Prokopicystis)'' {{Small|Cripps, 1989}}
*''[Proscotiaecystis](/source/Proscotiaecystis)'' {{Small|Ubaghs, 1994}}
*''[Protocystites](/source/Protocystites)'' {{Small|Hicks, 1872}}
*''[Reticulocarpos](/source/Reticulocarpos)'' {{Small|Jefferies & Prokop, 1972}}
*''[Scotiaecystis](/source/Scotiaecystis)'' {{Small|Caster & Ubaghs, 1967}}
*''[Thoralicystis](/source/Thoralicystis)'' {{Small|Chauvel, 1971}}
*''[Trigonocarpus](/source/Trigonocarpus)'' {{Small|Ubaghs, 1994}}
{{div col end}}

== See also ==
* [Cothurnocystidae](/source/Cothurnocystidae)

== References ==
{{Reflist}}
* {{cite journal|last1=Mus|first1=Monica Martí|title=The Ordovician cornute ''Flabellicystis rushtoni'' n. gen. n. sp. (Stylophora, Echinodermata) and its phylogenetic position within the group Cornuta|journal=Paläontologische Zeitschrift|date=April 2002|volume=76|issue=1|pages=99–116|doi=10.1007/BF02988189|s2cid=129282077 }}

== External links ==
* {{Wikispecies-inline}}
* {{Commons-inline}}
* [https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=31154 Cornuta at fossilworks.org] (retrieved 16 April 2016)

{{Taxonbar|from=Q21447200}}

Category:Homalozoa
Category:Prehistoric animal orders
Category:Echinoderm orders

{{paleo-echinoderm-stub}}

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Adapted from the Wikipedia article [Cornuta](https://en.wikipedia.org/wiki/Cornuta) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Cornuta?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.