# Cleorina

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***Cleorina*** is a genus of [leaf beetles](/source/Leaf_beetle) in the subfamily [Eumolpinae](/source/Eumolpinae). It is distributed in [South](/source/South_Asia), [East](/source/East_Asia) and [Southeast](/source/Southeast_Asia) [Asia](/source/Asia), [New Guinea](/source/New_Guinea) and [Australia](/source/Australia).

*[Cleorina modiglianii](/source/Cleorina_modiglianii)*, a species found in [Sumatra](/source/Sumatra), was considered as a potential [biological control agent](/source/Biological_control_agent) for the giant bramble (*[Rubus alceifolius](/source/Rubus_alceifolius)*), an invasive plant species in [La Réunion](/source/La_R%C3%A9union), but was rejected after testing its host specificity.[3]

## Species

Species include:[4][5][6][7][8][9]

- *[Cleorina aeneomicans](/source/Cleorina_aeneomicans)* ([Baly](/source/Joseph_Sugar_Baly), 1867)[10] – [Peninsular Malaysia](/source/Peninsular_Malaysia), [Java](/source/Java)
- *[Cleorina andamanensis](/source/Cleorina_andamanensis)* [Jacoby](/source/Martin_Jacoby), 1908[6] – [Andaman Islands](/source/Andaman_Islands)
- *[Cleorina assamensis](/source/Cleorina_assamensis)* [Jacoby](/source/Martin_Jacoby), 1908[6] – India ([Assam](/source/Assam), [Sikkim](/source/Sikkim), [Darjeeling district](/source/Darjeeling_district))
- *[Cleorina basalis](/source/Cleorina_basalis)* ([Baly](/source/Joseph_Sugar_Baly), 1867)[10] – Java
- *[Cleorina basipennis](/source/Cleorina_basipennis)* Medvedev, 1995[11] – Philippines ([Mindanao](/source/Mindanao))
- *[Cleorina bevani](/source/Cleorina_bevani)* ([Baly](/source/Joseph_Sugar_Baly), 1877) – India (Sikkim), Southern India
- *[Cleorina bicolor](/source/Cleorina_bicolor)* [Jacoby](/source/Martin_Jacoby), 1908[6] – Myanmar ([Tanintharyi Region](/source/Tanintharyi_Region))
- *[Cleorina bicoloripes](/source/Cleorina_bicoloripes)* [Pic](/source/Maurice_Pic), 1937 – Vietnam
- *[Cleorina borneoensis](/source/Cleorina_borneoensis)* Medvedev & Romantsov, 2014[12] – [Borneo](/source/Borneo) ([Sabah](/source/Sabah))
- *[Cleorina bosi](/source/Cleorina_bosi)* Medvedev, 2008[13] – [Sulawesi](/source/Sulawesi)
- *[Cleorina bryanti](/source/Cleorina_bryanti)* Medvedev, 2009[14] – Papua New Guinea
- *[Cleorina buechi](/source/Cleorina_buechi)* Medvedev, 2008[13] – Sulawesi
- *[Cleorina castanea](/source/Cleorina_castanea)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[1] – Philippines
- *[Cleorina chlorina](/source/Cleorina_chlorina)* Takizawa, 1988[15] – Nepal, India (Sikkim)
- *[Cleorina collaris](/source/Cleorina_collaris)* ([Baly](/source/Joseph_Sugar_Baly), 1867)[10] – Borneo ([Sarawak](/source/Sarawak))
- *[Cleorina costata](/source/Cleorina_costata)* Tan & Wang, 1981 – China ([Xizang](/source/Xizang))
- *[Cleorina costatella](/source/Cleorina_costatella)* Medvedev, 2009[16] – Vietnam
- *[Cleorina cyrtopus](/source/Cleorina_cyrtopus)* ([Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885) – Philippines
- *[Cleorina dohertyi](/source/Cleorina_dohertyi)* [Jacoby](/source/Martin_Jacoby), 1908[6] – Myanmar (Tanintharyi Region)
- *[Cleorina dura](/source/Cleorina_dura)* ([Weise](/source/Julius_Weise), 1923)[2] – Australia
- *[Cleorina femorata](/source/Cleorina_femorata)* Medvedev, 2009[14] – Western New Guinea
- *[Cleorina flavipes](/source/Cleorina_flavipes)* Medvedev, 2009[14] – Western New Guinea, Papua New Guinea
- *[Cleorina flavoornata](/source/Cleorina_flavoornata)* Medvedev, 1995[11] – Philippines ([Luzon](/source/Luzon))
- *[Cleorina fulva](/source/Cleorina_fulva)* [Jacoby](/source/Martin_Jacoby), 1908[6] – Southern India, Nepal
- *[Cleorina fulvicornis](/source/Cleorina_fulvicornis)* Medvedev, 2008[13] – Sulawesi
- *[Cleorina fulvipes](/source/Cleorina_fulvipes)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1890 – Sulawesi
- *[Cleorina fulvitarsis](/source/Cleorina_fulvitarsis)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1887 – Peninsular Malaysia, [Sumatra](/source/Sumatra)
- *[Cleorina gestroi](/source/Cleorina_gestroi)* [Jacoby](/source/Martin_Jacoby), 1896[17] – Sumatra
- *[Cleorina gorbunovi](/source/Cleorina_gorbunovi)* Medvedev, 2008[13] – Sulawesi
- *[Cleorina grandis](/source/Cleorina_grandis)* Eroshkina, 1988 – Vietnam
- *[Cleorina hainana](/source/Cleorina_hainana)* [Gressitt](/source/Judson_Linsley_Gressitt) & [Kimoto](/source/Shinsaku_Kimoto), 1961 – China ([Hainan](/source/Hainan)), Thailand, Laos, Vietnam
- *[Cleorina hirticollis](/source/Cleorina_hirticollis)* Bryant, 1950[18] – Western New Guinea, Papua New Guinea
- *[Cleorina imperialis](/source/Cleorina_imperialis)* ([Baly](/source/Joseph_Sugar_Baly), 1867)[10] – Peninsular Malaysia, Sumatra
- *[Cleorina indica](/source/Cleorina_indica)* [Jacoby](/source/Martin_Jacoby), 1908[6] – Myanmar (Tanintharyi Region)
- *[Cleorina instriata](/source/Cleorina_instriata)* [Pic](/source/Maurice_Pic), 1937 – China ([Yunnan](/source/Yunnan))
- *[Cleorina jacobyi](/source/Cleorina_jacobyi)* Duvivier, 1892 – India (Sikkim), Nepal
- *[Cleorina janthina](/source/Cleorina_janthina)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[1] – China, Nepal, Bhutan, India, Myanmar, Indochina
- *[Cleorina laeta](/source/Cleorina_laeta)* Medvedev, 1995[11] – Philippines (Luzon)
- *[Cleorina longicornica](/source/Cleorina_longicornica)* Tan, 1992 – China (Yunnan)
- *[Cleorina longicornis](/source/Cleorina_longicornis)* [Jacoby](/source/Martin_Jacoby), 1908[6] – Myanmar (Tanintharyi Region)
- *[Cleorina luzonica](/source/Cleorina_luzonica)* Medvedev, 1995[11] – Philippines (Luzon)
- *[Cleorina major](/source/Cleorina_major)* [Kimoto](/source/Shinsaku_Kimoto) & [Gressitt](/source/Judson_Linsley_Gressitt), 1982[5] – Laos
- *[Cleorina malayana](/source/Cleorina_malayana)* [Jacoby](/source/Martin_Jacoby), 1896[17] – Sumatra
- *[Cleorina manipurensis](/source/Cleorina_manipurensis)* [Jacoby](/source/Martin_Jacoby), 1908[6] – India ([Manipur](/source/Manipur))
- *[Cleorina metallica](/source/Cleorina_metallica)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[1] – Java
- *Cleorina metallica* Shukla, 1960 (homonym) – India ([Himachal Pradesh](/source/Himachal_Pradesh))
- *[Cleorina mimica](/source/Cleorina_mimica)* Medvedev & Eroshkina, 1983 – Vietnam
- *[Cleorina mjoebergi](/source/Cleorina_mjoebergi)* ([Weise](/source/Julius_Weise), 1923)[2] – Australia
- *[Cleorina modesta](/source/Cleorina_modesta)* [Jacoby](/source/Martin_Jacoby), 1908[6] – India ([Naga Hills](/source/Naga_Hills))
- *[Cleorina modiglianii](/source/Cleorina_modiglianii)* [Jacoby](/source/Martin_Jacoby), 1896[17] – Sumatra
- *[Cleorina morosa](/source/Cleorina_morosa)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[1] – Philippines
- *[Cleorina nepalensis](/source/Cleorina_nepalensis)* (Chûjô, 1966)[19] – India (Sikkim, Darjeeling district), Nepal
- *[Cleorina nigricornis](/source/Cleorina_nigricornis)* Medvedev, 2009[14] – Western New Guinea
- *[Cleorina nigrita](/source/Cleorina_nigrita)* [Jacoby](/source/Martin_Jacoby), 1895[20] – Borneo
- *[Cleorina nigroviridis](/source/Cleorina_nigroviridis)* [Jacoby](/source/Martin_Jacoby), 1908[6] – India (Assam)
- *[Cleorina nitidia](/source/Cleorina_nitidia)* Tan, 1981 – China (Xizang)
- *[Cleorina nitidicollis](/source/Cleorina_nitidicollis)* Tan & Wang, 1981 – China (Xizang)
- *[Cleorina nobilis](/source/Cleorina_nobilis)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[1] – India, Myanmar
- *[Cleorina oblonga](/source/Cleorina_oblonga)* [Jacoby](/source/Martin_Jacoby), 1908[6] – Myanmar ([Karen Hills](/source/Karen_Hills))
- *[Cleorina ornata](/source/Cleorina_ornata)* [Jacoby](/source/Martin_Jacoby), 1896[17] – Sumatra
- *[Cleorina papuana](/source/Cleorina_papuana)* Bryant, 1950[18] – Papua New Guinea
- *[Cleorina philippinensis](/source/Cleorina_philippinensis)* [Jacoby](/source/Martin_Jacoby), 1898 – Philippines
- *[Cleorina pulchella](/source/Cleorina_pulchella)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[1] – Australia
- *[Cleorina pulchra](/source/Cleorina_pulchra)* Medvedev, 2009[14] – Western New Guinea
- *[Cleorina puncticollis](/source/Cleorina_puncticollis)* [Jacoby](/source/Martin_Jacoby), 1894[21] – Peninsular Malaysia
- *[Cleorina punctipleuris](/source/Cleorina_punctipleuris)* Medvedev, 2008[13] – Sulawesi
- *[Cleorina punctisterna](/source/Cleorina_punctisterna)* Medvedev & Eroshkina, 1983 – Vietnam
- *[Cleorina purpurea](/source/Cleorina_purpurea)* [Lea](/source/Arthur_Mills_Lea), 1915[22] – Australia
- *[Cleorina purpureipennis](/source/Cleorina_purpureipennis)* ([Baly](/source/Joseph_Sugar_Baly), 1867)[10] – Peninsular Malaysia
- *[Cleorina riedeli](/source/Cleorina_riedeli)* Medvedev, 2009[14] – Papua New Guinea
- *[Cleorina robusta](/source/Cleorina_robusta)* Takizawa & Basu, 1987[23] – India (Sikkim)
- *[Cleorina schawalleri](/source/Cleorina_schawalleri)* Medvedev, 2009[14] – Papua New Guinea
- *[Cleorina sculpturata](/source/Cleorina_sculpturata)* ([Motschulsky](/source/Victor_Motschulsky), 1860)[24] – Myanmar
- *[Cleorina semipurpurea](/source/Cleorina_semipurpurea)* [Jacoby](/source/Martin_Jacoby), 1905 – Papua New Guinea (Ighibirei)
- *[Cleorina splendida](/source/Cleorina_splendida)* Bryant, 1950[18] – Papua New Guinea ([Kokoda](/source/Kokoda))
- *Cleorina splendida* Tan, 1992 (homonym) – China (Yunnan)
- *[Cleorina strigicollis](/source/Cleorina_strigicollis)* Medvedev & Eroshkina, 1983 – Vietnam
- *[Cleorina strigosipleuris](/source/Cleorina_strigosipleuris)* Medvedev, 2015[25] – China (Yunnan)
- *[Cleorina subnodosa](/source/Cleorina_subnodosa)* [Gressitt](/source/Judson_Linsley_Gressitt) & [Kimoto](/source/Shinsaku_Kimoto), 1961 – China (Hainan)
- *[Cleorina substriata](/source/Cleorina_substriata)* Medvedev, 2016[26] – Peninsular Malaysia
- *[Cleorina sulawensis](/source/Cleorina_sulawensis)* Medvedev, 2008[13] – Sulawesi
- *[Cleorina sumatrana](/source/Cleorina_sumatrana)* [Jacoby](/source/Martin_Jacoby), 1899 – Peninsular Malaysia, Sumatra
- *[Cleorina sumatrensis](/source/Cleorina_sumatrensis)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[1] – Sumatra
- *[Cleorina suturata](/source/Cleorina_suturata)* [Jacoby](/source/Martin_Jacoby), 1899 – [Bengal](/source/Bengal)
- *[Cleorina takizawai](/source/Cleorina_takizawai)* Medvedev & Sprecher-Uebersax, 1999[19] – India (Sikkim, Darjeeling district), Nepal
- *[Cleorina tibialis](/source/Cleorina_tibialis)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[1] – Philippines
- *[Cleorina verrucosa](/source/Cleorina_verrucosa)* Medvedev, 2008[13] – Sulawesi
- *[Cleorina vietnamica](/source/Cleorina_vietnamica)* Medvedev & Eroshkina, 1983 – Vietnam
- *[Cleorina viridis](/source/Cleorina_viridis)* ([Baly](/source/Joseph_Sugar_Baly), 1867)[10] – Sumatra
- *[Cleorina viridissima](/source/Cleorina_viridissima)* [Jacoby](/source/Martin_Jacoby), 1905 – Western New Guinea, Papua New Guinea
- *[Cleorina xizangensis](/source/Cleorina_xizangensis)* Tan & Wang, 1981 – China (Xizang)

Synonyms:

- *Cleorina bella* ([Jacoby](/source/Martin_Jacoby), 1892): synonym of *[Cleorina janthina](/source/Cleorina_janthina)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[9]
- *Cleorina bhamoensis* ([Jacoby](/source/Martin_Jacoby), 1892): synonym of *[Cleorina sculpturata](/source/Cleorina_sculpturata)* ([Motschulsky](/source/Victor_Motschulsky), 1860)[24]
- *Cleorina capitata* ([Jacoby](/source/Martin_Jacoby), 1892): synonym of *[Cleorina sculpturata](/source/Cleorina_sculpturata)* ([Motschulsky](/source/Victor_Motschulsky), 1860)[24]
- *Cleorina lefevrei* [Jacoby](/source/Martin_Jacoby), 1890: synonym of *[Cleorina janthina](/source/Cleorina_janthina)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[9]
- *Cleorina purpureipennis* var. *purpureicollis* [Pic](/source/Maurice_Pic), 1937: synonym of *[Cleorina janthina](/source/Cleorina_janthina)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[9]
- *Cleorina purpureipennis* var. *violaceipennis* [Pic](/source/Maurice_Pic), 1937: synonym of *[Cleorina janthina](/source/Cleorina_janthina)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[9]

Renamed species:

- *Cleorina costata* Medvedev & Eroshkina, 1983: renamed to *[Cleorina costatella](/source/Cleorina_costatella)* Medvedev, 2009[16]
- *Cleorina nepalensis* Takizawa, 1985 (preoccupied by *[Cleorina nepalensis](/source/Cleorina_nepalensis)* (Chûjô, 1966)): renamed to *[Cleorina takizawai](/source/Cleorina_takizawai)* Medvedev & Sprecher-Uebersax, 1999[19]

## References

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1. Weise, J. (1923). ["Chrysomeliden und Coccinelliden aus Queensland"](https://www.zin.ru/animalia/coleoptera/addpages/Andrey_Ukrainsky_Library/References_files/Weise23a.pdf). *Arkiv för Zoologi*. **15** (12): 1–150.

1. Le Bourgeois, T.; Baret, S.; de Chenon, R. D. (September 11–16, 2011). [*Biological Control of*Rubus alceifolius*(Rosaceae) in La Réunion Island (Indian Ocean): From Investigations on the Plant to the Release of the Biological Control Agent*Cibdela janthina*(Argidae)*](https://www.invasive.org/proceedings/pdfs/Le%20Bourgeois.pdf). Proceedings of the XIII International Symposium on Biological Control of Weeds. pp. 153–160.

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1. Kimoto, S. (1985). "Check-list of Chrysomelidae of South East Asia, South of Thailand and West of Irian-Jaya of Indonesia, III. Eumolpinae, 1". *Kurume University Journal*. **34** (1): 57–86.

1. Clavareau, H. (1914). ["Chrysomelidae: 11. Eumolpinae"](https://biodiversitylibrary.org/page/10432889). *Coleopterorum Catalogus*. Vol. 59. Junk, W. & Schenkling, S. (eds.). Berlin: [W. Junk](/source/Wilhelm_Junk). pp. 1–215.

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1. Baly, J. S. (1867). ["Phytophaga Malayana; a revision of the phytophagous beetles of the Malay Archipelago, with descriptions of the new species collected by Mr. A. R. Wallace"](https://biodiversitylibrary.org/page/14694386). *[Transactions of the Entomological Society of London](/source/Transactions_of_the_Entomological_Society_of_London)*. **4**: 1–300. 3. [doi:10.1111/j.1365-2311.1866.tb01857.x](https://doi.org/10.1111/j.1365-2311.1866.tb01857.x)

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1. Medvedev, L. & Romantsov, P. (2014). ["New and poorly known Chrysomelidae (Coleoptera) from Borneo"](https://web.archive.org/web/20170406192122/http://www.naturkundemuseum-bw.de/sites/default/files/publikationen/serie-a/ans07-12.pdf). *Stuttgarter Beiträge zur Naturkunde*. **7**: 235–251. A, Neue Serie. Archived from [the original](http://www.naturkundemuseum-bw.de/sites/default/files/publikationen/serie-a/ans07-12.pdf) on 2017-04-06. Retrieved 2018-08-05.

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1. Medvedev, Lev N. (2009). ["New genera and species of Zeugophorinae and Eumolpinae (Coleoptera: Chrysomelidae) from New Guinea"](https://web.archive.org/web/20130830232644/http://www.naturkundemuseum-bw.de/sites/default/files/publikationen/serie-a/ans02-11medvedev.pdf). *Stuttgarter Beiträge zur Naturkunde A, Neue Serie*. **2**: 371–408. Archived from [the original](http://www.naturkundemuseum-bw.de/sites/default/files/publikationen/serie-a/ans02-11medvedev.pdf) on 2013-08-30. Retrieved 2018-08-05.

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1. Medvedev, L. N. (2000). ["Jacoby's types of Chrysomelidae (Coleoptera) from Burma in the Museo Civico di Storia Naturale "Giacomo Doria", Genoa. Part 1"](https://www.biodiversitylibrary.org/page/63848626). *Annali del Museo Civico di Storia Naturale "Giacomo Doria"*. **93**: 167–184.

1. Medvedev, L. N. (2015). ["New and poorly known Oriental Chrysomelidae (Insecta: Coleoptera) in the collection of the Naturkundemuseum Erfurt"](https://www.naturkundemuseum-erfurt.de/fileadmin/user_upload/Wissenschaft/Publikationen/Museumshefte_Vernate/Hefte/34/21_medvedev_1.pdf). *Vernate*. **34**: 319–335.

1. Medvedev, L. N. (2016). ["New and poorly known Oriental Chrysomelidae (Insecta: Coleoptera) in the collection of the Naturkundemuseum Erfurt"](https://www.naturkundemuseum-erfurt.de/fileadmin/user_upload/Wissenschaft/Publikationen/Museumshefte_Vernate/Hefte/35/15_medvedev.pdf). *Vernate*. **35**: 347–365.

## External links

- [Genus *Cleorina* Lefèvre, 1885](https://biodiversity.org.au/afd/taxa/Cleorina) at [Australian Faunal Directory](/source/Australian_Faunal_Directory)

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Adapted from the Wikipedia article [Cleorina](https://en.wikipedia.org/wiki/Cleorina) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Cleorina?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.