# Cleoporus

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***Cleoporus*** is a genus of [leaf beetles](/source/Leaf_beetle) in the subfamily [Eumolpinae](/source/Eumolpinae). It is distributed in [South](/source/South_Asia), [East](/source/East_Asia) and [Southeast](/source/Southeast_Asia) [Asia](/source/Asia).[2][3][4]

## Species

- *[Cleoporus aedilis](/source/Cleoporus_aedilis)* ([Weise](/source/Julius_Weise), 1922)[5][6] – Philippines ([Luzon](/source/Luzon))
- *[Cleoporus aeneipennis](/source/Cleoporus_aeneipennis)* [Chen](/source/Chen_Shixiang), 1935 – Vietnam
- *[Cleoporus aenescens](/source/Cleoporus_aenescens)* ([Jacoby](/source/Martin_Jacoby), 1908)[4][7] – Myanmar
- *[Cleoporus badius](/source/Cleoporus_badius)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1889 – Thailand, Cambodia, Laos, Vietnam, China ([Guangdong](/source/Guangdong), [Guangxi](/source/Guangxi), [Hainan](/source/Hainan), [Hong Kong](/source/Hong_Kong), [Hunan](/source/Hunan))
- *[Cleoporus costatus](/source/Cleoporus_costatus)* Chûjô, 1956[8] – Taiwan
- *[Cleoporus cuprescens](/source/Cleoporus_cuprescens)* ([Baly](/source/Joseph_Sugar_Baly), 1878)[9][10] (doubtful assignment) – Pakistan, India
- *[Cleoporus dalatensis](/source/Cleoporus_dalatensis)* Medvedev, 1998[11] – Vietnam
- *[Cleoporus harmandi](/source/Cleoporus_harmandi)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1893[12] – Vietnam
- *[Cleoporus inornatus](/source/Cleoporus_inornatus)* [Jacoby](/source/Martin_Jacoby), 1908[4] – India, Sri Lanka
- *[Cleoporus laevifrons](/source/Cleoporus_laevifrons)* ([Jacoby](/source/Martin_Jacoby), 1908)[4][7] – Myanmar
- *[Cleoporus lateralis](/source/Cleoporus_lateralis)* ([Motschulsky](/source/Victor_Motschulsky), 1866)[13] – India, Myanmar
- *[Cleoporus lefevrei](/source/Cleoporus_lefevrei)* Duvivier, 1892 – [Bengal](/source/Bengal)
- *[Cleoporus lineatus](/source/Cleoporus_lineatus)* Medvedev & Sprecher-Uebersax, 1999[14] – Nepal
- *[Cleoporus maculicollis](/source/Cleoporus_maculicollis)* [Jacoby](/source/Martin_Jacoby), 1904 – India ([Khandesh](/source/Khandesh))
- *[Cleoporus pallidipes](/source/Cleoporus_pallidipes)* ([Fairmaire](/source/L%C3%A9on_Fairmaire), 1888) – China ([Jiangxi](/source/Jiangxi))
- *[Cleoporus plagiatus](/source/Cleoporus_plagiatus)* [Jacoby](/source/Martin_Jacoby), 1892 – Myanmar
- *[Cleoporus quadripustulatus](/source/Cleoporus_quadripustulatus)* ([Baly](/source/Joseph_Sugar_Baly), 1859)[15][6] – Philippines ([Mindanao](/source/Mindanao))
- *[Cleoporus recticollis](/source/Cleoporus_recticollis)* ([Pic](/source/Maurice_Pic), 1929)[7] – North Vietnam
- *[Cleoporus robustus](/source/Cleoporus_robustus)* ([Baly](/source/Joseph_Sugar_Baly), 1874)[16] – East and South China, South Korea, Japan
- *[Cleoporus similis](/source/Cleoporus_similis)* Medvedev & Eroshkina, 1985 – Vietnam
- *[Cleoporus sumbaensis](/source/Cleoporus_sumbaensis)* [Jacoby](/source/Martin_Jacoby), 1899[17] – [Sumba](/source/Sumba)
- *[Cleoporus suturalis](/source/Cleoporus_suturalis)* [Chen](/source/Chen_Shixiang), 1935 – China (Guangdong, Hainan), Vietnam
- *[Cleoporus taynguensis](/source/Cleoporus_taynguensis)* Medvedev & Eroshkina, 1985 – Vietnam
- *[Cleoporus tibialis](/source/Cleoporus_tibialis)* ([Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885) – Vietnam
- *[Cleoporus timorensis](/source/Cleoporus_timorensis)* [Jacoby](/source/Martin_Jacoby), 1894[18] – [Timor](/source/Timor)
- *[Cleoporus trimaculatus](/source/Cleoporus_trimaculatus)* [Kimoto](/source/Shinsaku_Kimoto) & [Gressitt](/source/Judson_Linsley_Gressitt), 1982[2] – Thailand
- *[Cleoporus udovichenkoi](/source/Cleoporus_udovichenkoi)* Medvedev & Romantsov, 2013[19] – South Vietnam
- *[Cleoporus variabilis](/source/Cleoporus_variabilis)* ([Baly](/source/Joseph_Sugar_Baly), 1874)[16] – Russia (southern Far East), North-Eastern China, Taiwan, North Korea, Japan
- *[Cleoporus vietnamensis](/source/Cleoporus_vietnamensis)* Medvedev & Eroshkina, 1985 – Vietnam

The following are synonyms of other species:[20]

- *Cleoporus birmanicus* [Jacoby](/source/Martin_Jacoby), 1892: synonym of *[Cleoporus lateralis](/source/Cleoporus_lateralis)* ([Motschulsky](/source/Victor_Motschulsky), 1866)
- *Cleoporus cruciatus* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1884:[1] synonym of *[Cleoporus quadripustulatus](/source/Cleoporus_quadripustulatus)* ([Baly](/source/Joseph_Sugar_Baly), 1859)[6]
- *Cleoporus jacobyi* Medvedev & Eroshkina, 1985 (replacement name for *Cleoporus lefevrei* [Jacoby](/source/Martin_Jacoby), 1895 nec Duvivier, 1892): synonym of *[Cleoporus inornatus](/source/Cleoporus_inornatus)* [Jacoby](/source/Martin_Jacoby), 1908
- *Cleoporus lefevrei* [Jacoby](/source/Martin_Jacoby), 1895 nec Duvivier, 1892: synonym of *[Cleoporus inornatus](/source/Cleoporus_inornatus)* [Jacoby](/source/Martin_Jacoby), 1908
- *Cleoporus variegatus* [Jacoby](/source/Martin_Jacoby), 1904: transferred to *[Bathseba](/source/Bathseba_(beetle))*[21]

## References

1. Lefèvre, E. (1884). ["\[*Cleoporus*\]"](https://biodiversitylibrary.org/page/32550560). *[Bulletin de la Société Entomologique de France](/source/Bulletin_de_la_Soci%C3%A9t%C3%A9_Entomologique_de_France)*. **4**: LXXVI–LXXVII.

1. Kimoto, S. & Gressitt, J. L. (1982). ["Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos and Vietnam. III. Eumolpinae"](https://catalog.lib.kyushu-u.ac.jp/opac_download_md/2421/1.pdf). *Esakia*. **18**: 1–141. [doi:10.5109/2421](https://doi.org/10.5109/2421). [hdl:2324/2421](https://hdl.handle.net/2324/2421). [S2CID 83265328](https://api.semanticscholar.org/CorpusID:83265328)

1. Moseyko, A. G. & Sprecher-Uebersax, E. (2010). ["Eumolpinae"](https://books.google.com/books?id=qt8zDwAAQBAJ&pg=PA620). *Catalogue of Palaearctic Coleoptera. Volume 6. Chrysomeloidea*. Löbl, I. & Smetana, A. (eds.). Stenstrup, Denmark: Apollo Books. pp. 619–643. ISBN 978-87-88757-84-2.

1. Jacoby, M. (1908). [*Coleoptera. Chrysomelidae. Vol. 1*](https://archive.org/details/coleopterachryso00jacoiala). [The Fauna of British India, Including Ceylon and Burma](/source/The_Fauna_of_British_India,_Including_Ceylon_and_Burma). Bingham, C. T. (ed.). London: [Taylor & Francis](/source/Taylor_%26_Francis)

1. Weise, J. (1922). ["Chrysomeliden der Philippinen: III"](https://web.archive.org/web/20210515233228/https://philjournalsci.dost.gov.ph/images/pdf_upload/pjs1922/PJS_Vol_21_No5_Nov_1922.pdf). *The Philippine Journal of Science*. **21** (5): 423–490. Archived from [the original](http://philjournalsci.dost.gov.ph/images/pdf_upload/pjs1922/PJS_Vol_21_No5_Nov_1922.pdf) on May 15, 2021.

1. Moseyko, A. G. (2012 [2011]). "To the knowledge of the leaf-beetle tribe Nodinini (Coleoptera, Chrysomelidae, Eumolpinae) from the Philippines and Borneo". *Entomological Review*. **92** (3): 315–328. [doi:10.1134/S0013873812030086](https://doi.org/10.1134/S0013873812030086). [S2CID 15106058](https://api.semanticscholar.org/CorpusID:15106058)

1. Moseyko, A. G. (2013 [2012]). "On the distribution and synonymy of some indo-malayan species of the chrysomelid genus *Pagria* Lefèvre, 1884 (Coleoptera, Chrysomelidae: Eumolpinae)". *Entomological Review*. **93** (2): 208–213. [doi:10.1134/S0013873813020097](https://doi.org/10.1134/S0013873813020097). [S2CID 15837259](https://api.semanticscholar.org/CorpusID:15837259)

1. Chûjô, M. (1956). ["A taxonomic study on the Chrysomelidae (Insecta: Coleoptera) from Formosa. Part VIII. Subfamily Eumolpinae"](https://web.archive.org/web/20210515232250/https://philjournalsci.dost.gov.ph/images/pdf_upload/pjs1956/PJS_Vol_85_No1_Mar_1956.pdf). *The Philippine Journal of Science*. **85** (1): 1–180. Archived from [the original](http://philjournalsci.dost.gov.ph/images/pdf_upload/pjs1956/PJS_Vol_85_No1_Mar_1956.pdf) on May 15, 2021.

1. Baly, J. S. (1878). ["Descriptions of the phytophagous Coleoptera collected by the late Dr. F. Stoliczka during Forsyth's expedition to Kashgar in 1873-74"](https://biodiversitylibrary.org/page/10891031). *Cistula Entomologica*. **2**: 369–383.

1. Moseyko, A. G. & Sprecher-Uebersax, E. (2010). ["New nomenclatural and taxonomic acts, and comments. Chrysomelidae: Eumolpinae"](https://books.google.com/books?id=qt8zDwAAQBAJ&pg=PA80). *Catalogue of Palaearctic Coleoptera. Volume 6. Chrysomeloidea*. Löbl, I. & Smetana, A. (eds.). Stenstrup, Denmark: Apollo Books. pp. 80–83. ISBN 978-87-88757-84-2.

1. Medvedev, L.N. (1998). ["New Chrysomelidae (Coleoptera) from Southeast Asia in the Hungarian Natural History Museum"](http://publication.nhmus.hu/pdf/annHNHM/Annals_HNHM_1998_Vol_90_163.pdf). *Annales Historico-Naturales Musei Nationalis Hungarici*. **90**: 163–174.

1. Lefèvre, E. (1893). ["Contributions à la faune Indo-Chinoise, 12. Clytrides & Eumolpides"](https://biodiversitylibrary.org/page/8236945). *[Annales de la Société Entomologique de France](/source/Annales_de_la_Soci%C3%A9t%C3%A9_Entomologique_de_France)*. **62**: 111–134.

1. Medvedev, L. N. (2007). ["To the knowledge of Chrysomelidae (Coleoptera) described by V. Motschulsky"](http://zmmu.msu.ru/files/images/spec/Russ%20Ent%20J/REJ_15-4_409-417%20(Medvedev).pdf). *Russian Entomological Journal*. **15** (4): 409–417.

1. Medvedev, L. N. & Sprecher-Uebersax, E. (1999). "Taxonomical study of Chrysomelidae (Coleoptera) from Nepal". *Entomologica Basiliensia*. **21**: 355–370. [doi:10.5169/seals-980426](https://doi.org/10.5169/seals-980426)

1. Baly, J. S. (1859). "Descriptions of new genera and species of Phytophagous insects". *[Annals and Magazine of Natural History](/source/Annals_and_Magazine_of_Natural_History)*. **4** (20): 124–128. 3. [doi:10.1080/00222935908697096](https://doi.org/10.1080/00222935908697096)

1. Baly, J. S. (1874). ["Catalogue of the Phytophagous Coleoptera of Japan, with descriptions of the species new to science"](https://www.biodiversitylibrary.org/part/27159). *[Transactions of the Entomological Society of London](/source/Transactions_of_the_Entomological_Society_of_London)*. **22** (2): 161–217. [doi:10.1111/j.1365-2311.1874.tb00164.x](https://doi.org/10.1111/j.1365-2311.1874.tb00164.x)

1. Jacoby, M. (1899). ["Descriptions of the new species of phytophagous Coleoptera obtained by Dr. Dohrn in Sumatra"](https://www.zobodat.at/pdf/Entomologische-Zeitung-Stettin_60_0259-0313.pdf). *Stettiner Entomologische Zeitung*. **60**: 259–313, 1 pl.

1. Jacoby, M. (1894). ["Descriptions of new genera and species of phytophagous Coleoptera obtained by W. Doherty in the Malayan Archipelago"](https://www.biodiversitylibrary.org/page/3857542). *[Novitates Zoologicae](/source/Novitates_Zoologicae)*. **1**: 267–330. [doi:10.5962/bhl.part.24565](https://doi.org/10.5962/bhl.part.24565)

1. Medvedev, L.N. & Romantsov, P.V. (2013). ["New and poorly known Chrysomelidae (Coleoptera) from South-East Asia"](http://ssc-ras.ru/files/files/25_%20Medvedev.pdf). *Caucasian Entomological Bulletin*. **9** (1): 137–140. [doi:10.23885/1814-3326-2013-9-1-137-140](https://doi.org/10.23885/1814-3326-2013-9-1-137-140)

1. Moseyko, A.G. (2020). "Notes on Asiatic Eumolpinae (Coleoptera, Chrysomelidae)". *Entomological Review*. **100** (6): 843–862. [doi:10.1134/S0013873820060123](https://doi.org/10.1134/S0013873820060123). [S2CID 234621478](https://api.semanticscholar.org/CorpusID:234621478)

1. Kumari, S.A.; Moseyko, A.G.; Prathapan, K.D. (2023). ["A new synonym of *Bathseba* Motschulsky, 1866 (Chrysomelidae, Eumolpinae, Typophorini) with a new species and redescription of *B. bifasciata* (Jacoby, 1895) n. comb. and other taxonomic and biological notes"](https://tb.plazi.org/GgServer/summary/E2566F2CB523FFA4FFC5FF89963CFFB1). *[Zootaxa](/source/Zootaxa)*. **5293** (1): 122–144. [doi:10.11646/zootaxa.5293.1.5](https://doi.org/10.11646/zootaxa.5293.1.5). [PMID 37518494](https://pubmed.ncbi.nlm.nih.gov/37518494). [S2CID 258810165](https://api.semanticscholar.org/CorpusID:258810165)

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Adapted from the Wikipedia article [Cleoporus](https://en.wikipedia.org/wiki/Cleoporus) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Cleoporus?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.