# Cingulata

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> Markdown URL: https://mediated.wiki/source/Cingulata.md
> Source: https://en.wikipedia.org/wiki/Cingulata
> Source revision: 1350696583
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{{Short description|Order of armored mammals from the Americas}}
{{Automatic taxobox
| fossil_range = [Late Paleocene](/source/Late_Paleocene)-Recent, {{Fossil range|58.7|0}}
| image = Cingulata2.jpg
| image_caption = ''[Glyptodon](/source/Glyptodon)'' ([Vienna](/source/Natural_History_Museum%2C_Vienna)) and ''[Dasypus novemcinctus](/source/Dasypus_novemcinctus)''
| taxon = Cingulata
| authority = [Illiger](/source/Johann_Karl_Wilhelm_Illiger), 1811
| subdivision_ranks = Families
| subdivision = * [Chlamyphoridae](/source/Chlamyphoridae)
* [Dasypodidae](/source/Dasypodidae)
* [†](/source/Extinct)[Pachyarmatheriidae](/source/Pachyarmatheriidae)
* [†](/source/Extinct)[Pampatheriidae](/source/Pampatheriidae)
* [†](/source/Extinct)[Peltephilidae](/source/Peltephilidae)
| range_map = Cingulata range.jpg
| range_map2 = Armadillo species density.jpg
| range_map2_caption = The distribution and density of Cingulata species
}}

'''Cingulata''', part of the superorder [Xenarthra](/source/Xenarthra), is an [order](/source/order_(biology)) of [armored](/source/armour_(zoology)) [New World](/source/Americas) [placental](/source/placental) [mammal](/source/mammal)s. The [armadillo](/source/armadillo)s, whose species are split between the families [Dasypodidae](/source/Dasypodidae) and [Chlamyphoridae](/source/Chlamyphoridae), are the only surviving members of the order.<ref name = MSW3>{{MSW3 Cingulata | id = 11700001}}</ref> Two groups of cingulates much larger than extant armadillos (maximum body mass of 45&nbsp;kg (100&nbsp;lb) in the case of the giant armadillo<ref>[http://www.faunaparaguay.com/mamm6Priodontesmaximus.pdf Giant Armadillo ''Priodontes maximus'' (Kerr, 1792)]. FaunaParaguay.com</ref>) existed until recently: [pampatheriids](/source/Pampatheriidae), which reached weights of up to 200&nbsp;kg (440&nbsp;lb)<ref name = "Vizcaíno">{{cite journal
  | last = Vizcaíno | first = S. F. |author2=De Iuliis, G. |author3=Bargo, M. S. | title = Skull Shape, Masticatory Apparatus, and Diet of ''Vassallia'' and ''Holmesina'' (Mammalia: Xenarthra: Pampatheriidae): When Anatomy Constrains Destiny | journal = Journal of Mammalian Evolution | volume = 5 | issue = 4 | pages = 291–322 | year = 1998
 | doi = 10.1023/A:1020500127041 | s2cid = 20186439 }}</ref> and chlamyphorid [glyptodont](/source/glyptodont)s, which attained masses of 2,000&nbsp;kg (4,400&nbsp;lb)<ref name="Blanco2009">{{cite journal|last1=Blanco|first1=R. E.|last2=Jones|first2=W. W.|last3=Rinderknecht|first3=A.|title=The sweet spot of a biological hammer: the centre of percussion of glyptodont (Mammalia: Xenarthra) tail clubs|journal=Proceedings of the Royal Society B: Biological Sciences|volume= 276|issue=1675|date= 2009-08-26|pages= 3971–3978|issn=0962-8452|doi=10.1098/rspb.2009.1144|pmid=19710060|pmc=2825778}}</ref> or more.

The cingulate order originated in [South America](/source/South_America) during the [Paleocene epoch](/source/Paleocene_epoch) about 66 to 56&nbsp;million years ago, and due to the continent's former isolation remained confined to it during most of the [Cenozoic](/source/Cenozoic). However, the formation of a [land bridge](/source/Isthmus_of_Panama) allowed members of all three families to migrate to southern North America during the [Pliocene](/source/Pliocene)<ref name = "Mead_2007">{{cite journal
 | last = Mead | first = J. I. |author2=Swift, S. L. |author3=White, R. S. |author4=McDonald, H. G. |author5=Baez, A.
 | title = Late Pleistocene (Rancholabrean) Glyptodont and Pampathere (Xenarthra, Cingulata) from Sonora, Mexico
 | journal = [Revista Mexicana de Ciencias Geológicas](/source/Revista_Mexicana_de_Ciencias_Geol%C3%B3gicas)
 | volume = 24 | issue = 3 | pages = 439–449 (see p. 440)
 | year = 2007
 | url = http://www.scielo.org.mx/pdf/rmcg/v24n3/v24n3a11.pdf 
 | access-date = 2013-06-15}}</ref> or early [Pleistocene](/source/Pleistocene)<ref name="Woodburne2010">{{cite journal|last1=Woodburne|first1=M. O.|title=The Great American Biotic Interchange: Dispersals, Tectonics, Climate, Sea Level and Holding Pens|journal=Journal of Mammalian Evolution|volume= 17|issue= 4|date= 2010-07-14|pages= 245–264 (see p. 249)|issn= 1064-7554|doi= 10.1007/s10914-010-9144-8|pmid=21125025|pmc=2987556}}</ref> as part of the [Great American Interchange](/source/Great_American_Interchange). After surviving for tens of millions of years, all of the pampatheriids and giant [glyptodont](/source/glyptodont)s apparently died out during the [Quaternary extinction event](/source/Quaternary_extinction_event) at the beginning of the [Holocene](/source/Holocene),<ref name="HubbeHubbe2013">{{cite journal|last1=Hubbe|first1=A.|last2=Hubbe|first2=M.|last3=Neves|first3=W. A.|title=The Brazilian megamastofauna of the Pleistocene/Holocene transition and its relationship with the early human settlement of the continent|journal=Earth-Science Reviews|volume=118|date=March 2013|pages=1–10 (see pages 3, 6)|issn=0012-8252|doi=10.1016/j.earscirev.2013.01.003|bibcode=2013ESRv..118....1H}}</ref><ref name = "Fiedal">{{Cite book
  | first = Stuart | last = Fiedal | editor-last = Haynes | editor-first = Gary
  | contribution = Sudden Deaths: The Chronology of Terminal Pleistocene Megafaunal Extinction
  | title = American Megafaunal Extinctions at the End of the Pleistocene
  | year = 2009 | pages = 21–37 (see p. 31)| publisher = [Springer](/source/Springer_Science%2BBusiness_Media)
  | doi = 10.1007/978-1-4020-8793-6_2 | isbn = 978-1-4020-8792-9 | oclc = 313368423| series = Vertebrate Paleobiology and Paleoanthropology }}</ref> along with much of the rest of the regional [megafauna](/source/megafauna), shortly after the colonization of the Americas by [Paleo-Indians](/source/Paleo-Indians).

==Description==
Armadillos have dorsal armor that is formed by [osteoderm](/source/osteoderm)s, plates of dermal bone covered in relatively small, overlapping [keratin](/source/keratin)ized epidermal scales called "[scute](/source/scute)s". Most species have rigid shields over the shoulders and hips, with three to nine bands separated by flexible skin covering the back and flanks.<ref name="EoM">{{cite book |editor=Macdonald, D. |author=Dickman, Christopher R. |year=1984 |title=The Encyclopedia of Mammals |publisher=Facts on File |location=New York |pages=[https://archive.org/details/encyclopediaofma00mals_0/page/781 781–783] |isbn=978-0-87196-871-5 |url-access=registration |url=https://archive.org/details/encyclopediaofma00mals_0/page/781 }}</ref>

Pampatheres also had shells that were flexible due to three movable lateral bands of osteoderms.<ref name = "Vizcaíno"/> The osteoderms of pampatheres were each covered by a single scute, unlike those of armadillos, which have more than one.<ref name = "Vizcaíno"/> Glyptodonts, on the other hand, had rigid, turtle-like shells of fused osteoderms.

Both groups have or had a cap of armor atop their heads. Glyptodonts also had heavily armored tails; some, such as ''[Doedicurus](/source/Doedicurus)'', had mace-like clubs at the ends of their tails, similar to those of [ankylosaur](/source/ankylosaur)s, evidently used for defensive or [agonistic](/source/Agonistic_behaviour) purposes.<ref name="Blanco2009" />

Most armadillos [eat insects](/source/insectivore) and other invertebrates; some are more [omnivorous](/source/omnivorous) and may also eat small vertebrates and vegetable matter. Pampatheres are thought to have been specialized for [grazing](/source/grazing),<ref name = "Vizcaíno"/> and isotopic analysis indicates the diet of glyptodonts was dominated by [C4](/source/C4_carbon_fixation) [grass](/source/grass)es.<ref name="Pérez-CrespoArroyo-Cabrales2011">{{cite journal|last1=Pérez-Crespo|first1=V. A.|last2=Arroyo-Cabrales|first2=J.|last3=Alva-Valdivia|first3=L. M.|last4=Morales-Puente|first4=P.|last5=Cienfuegos-Alvarado|first5=E.|title=Diet and habitat definitions for Mexican glyptodonts from Cedral (San Luis Potosí, México) based on stable isotope analysis|journal=Geological Magazine|volume=149|issue=1|date=2011-10-18|pages=153–157|issn=0016-7568|doi=10.1017/S0016756811000951|s2cid=129862616}}</ref> [Euphractinae](/source/Euphractinae) is unique for speciations towards [carnivory](/source/carnivory), culminating in the macropredatory genus ''[Macroeuphractus](/source/Macroeuphractus)''.

==Classification==
{{main|List of cingulates}}
[[File:Gyptodon Cosmo Caixa.JPG|thumb|right|''[Holmesina](/source/Holmesina) septentrionalis'' ([Barcelona](/source/CosmoCaixa_Barcelona))]]
[[File:Nine-banded armadillo skeleton.jpg|thumb|right|[Nine-banded armadillo](/source/Nine-banded_armadillo), ''D.&nbsp;novemcinctus'' ([Smithsonian](/source/Smithsonian_National_Museum_of_Natural_History))]]
[[File:Glyptodon clavipes 01.jpg|thumb|right|''[Glyptodon](/source/Glyptodon) clavipes'' ([Berlin](/source/Natural_History_Museum%2C_Berlin))]]
thumb|right
The taxonomic table below follows the results of a [phylogenetic analysis](/source/phylogenetic_analysis) published by Delsuc ''et al.'', 2016. While glyptodonts have traditionally been considered stem-group cingulates outside the group that contains modern armadillos, this 2016 study conducted an analysis of ''[Doedicurus](/source/Doedicurus)'' [mtDNA](/source/mtDNA) and found that it was, in fact, nested within the modern armadillos as the [sister group](/source/sister_group) of a [clade](/source/clade) consisting of Chlamyphorinae and Tolypeutinae.<ref name=Delsuc2016>
{{cite journal
 |last1= Delsuc    |first1= F.     |last2= Gibb      |first2=G.C.
 |last3= Kuch      |first3= M.     |last4= Billet    |first4= G.
 |last5= Hautier   |first5= L.     |last6= Southon   |first6= J.
 |last7= Rouillard |first7= J.-M.  |last8= Fernicola |first8=J.C.
 |last9= Vizcaíno  |first9=S.F.    |last10= MacPhee  |first10=R.D.E.
 |last11= Poinar   |first11=H.N.     |display-authors=6
 |date= 2016-02-22  |df=dmy-all
 |title=The phylogenetic affinities of the extinct glyptodonts
 |journal= Current Biology
 |volume= 26 |issue =4 |pages= R155–R156
 |doi= 10.1016/j.cub.2016.01.039 |doi-access= free
 |pmid= 26906483 |hdl= 11336/49579 |hdl-access= free
 |url= https://hal.archives-ouvertes.fr/hal-01879335  |via=archives-ouvertes.fr
}}
</ref>

'''Order Cingulata'''
* '''Family''' [†](/source/Extinct)'''[Peltephilidae](/source/Peltephilidae)''' <small>Ameghino 1894</small>
** Genus [†](/source/Extinct)''[Anantiosodon](/source/Anantiosodon)'' <small>Ameghino 1891</small>
** Genus [†](/source/Extinct)''[Epipeltephilus](/source/Epipeltephilus)'' <small>Ameghino 1904</small>
** Genus [†](/source/Extinct)''[Parapeltecoelus](/source/Parapeltecoelus)'' <small>Bordas 1938</small>
** Genus [†](/source/Extinct)''[Peltecoelus](/source/Peltecoelus)'' <small>Ameghino 1902</small>
** Genus [†](/source/Extinct)''[Peltephilus](/source/Peltephilus)'' <small>Ameghino 1887</small> (Horned armadillo)
** Genus [†](/source/Extinct)''[Ronwolffia](/source/Ronwolffia)'' <small>Shockey 2017</small>
* '''Family''' [†](/source/Extinct)'''[Paleopeltidae](/source/Paleopeltidae)''' <small>Ameghino 1895</small>
** Genus [†](/source/Extinct)''[Palaeopeltis](/source/Palaeopeltis)'' <small>Ameghino 1895</small>
* '''Family''' [†](/source/Extinct)'''[Pampatheriidae](/source/Pampatheriidae)''' 
** Genus ?[†](/source/Extinct)''[Machlydotherium](/source/Machlydotherium)'' <small>Ameghino 1902</small>
** Genus [†](/source/Extinct)''[Holmesina](/source/Holmesina)'' <small>Simpson 1930</small>
** Genus [†](/source/Extinct)''[Kraglievichia](/source/Kraglievichia)'' <small>Castellanos 1927</small>
** Genus [†](/source/Extinct)''[Machlydotherium](/source/Machlydotherium)'' 
** Genus [†](/source/Extinct)''[Pampatherium](/source/Pampatherium)'' <small>Ameghino 1875 ex Gervais & Ameghino 1880</small>
** Genus [†](/source/Extinct)''[Scirrotherium](/source/Scirrotherium)'' <small>Edmund & Theodor 1997</small>
** Genus [†](/source/Extinct)''[Tonnicinctus](/source/Tonnicinctus)''  <small>Góis et al. 2015</small><ref>{{Cite journal|author1=Flávio Góis |author2=Laureano Raúl González Ruiz |author3=Gustavo Juan Scillato-Yané |author4=Esteban Soibelzon |year=2015 |title=A Peculiar New Pampatheriidae (Mammalia: Xenarthra: Cingulata) from the Pleistocene of Argentina and Comments on Pampatheriidae Diversity |journal=PLOS ONE |volume=10 |issue=6 |article-number=e0128296 |doi=10.1371/journal.pone.0128296 |pmid=26083486 |pmc=4470999|bibcode=2015PLoSO..1028296G |doi-access=free }}</ref>
** Genus [†](/source/Extinct)''[Vassallia](/source/Vassallia)'' <small>Castellanos 1927</small> [''[Plaina](/source/Plaina)'' <small>Castellanos 1937</small>]
** Genus [†](/source/Extinct)''[Yuruatherium](/source/Yuruatherium)'' <small>Ciancio et al. 2012</small>
* '''Family''' [†](/source/Extinct)'''[Pachyarmatheriidae](/source/Pachyarmatheriidae)''' <small>Fernicola et al. 2018</small>
** Genus [†](/source/Extinct)''[Neoglyptatelus](/source/Neoglyptatelus)'' <small>Carlini, Vizcaíno & Scillato-Yané 1997</small>
** Genus [†](/source/Extinct)''[Pachyarmatherium](/source/Pachyarmatherium)'' <small>Downing & White 1995</small>
* '''Family [Dasypodidae](/source/Dasypodidae)''' (long-nosed armadillos)
** Genus [†](/source/Extinct)''[Acantharodeia](/source/Acantharodeia)''
** Genus [†](/source/Extinct)''[Amblytatus](/source/Amblytatus)''
** Genus [†](/source/Extinct)''[Archaeutatus](/source/Archaeutatus)''
** Genus [†](/source/Extinct)''[Astegotherium](/source/Astegotherium)''
** Genus [†](/source/Extinct)''[Barrancatatus](/source/Barrancatatus)''
** Genus [†](/source/Extinct)''[Chasicotatus](/source/Chasicotatus)''
** Genus [†](/source/Extinct)''[Chorobates](/source/Chorobates_(genus))''
** Genus [†](/source/Extinct)''[Coelutaetus](/source/Coelutaetus)''
** Genus [†](/source/Extinct)''[Eocoleophorus](/source/Eocoleophorus)''
** Genus [†](/source/Extinct)''[Epipeltecoelus](/source/Epipeltecoelus)''
** Genus [†](/source/Extinct)''[Eutatus](/source/Eutatus)''
** Genus [†](/source/Extinct)''[Hemiutaetus](/source/Hemiutaetus)''
** Genus [†](/source/Extinct)''[Isutaetus](/source/Isutaetus)''
** Genus [†](/source/Extinct)''[Lumbreratherium](/source/Lumbreratherium)''
** Genus [†](/source/Extinct)''[Macrochorobates](/source/Macrochorobates)''
** Genus [†](/source/Extinct)''[Mazzoniphractus](/source/Mazzoniphractus)''
** Genus [†](/source/Extinct)''[Meteutatus](/source/Meteutatus)''
** Genus [†](/source/Extinct)''[Pedrolypeutes](/source/Pedrolypeutes)''
** Genus [†](/source/Extinct)''[Prodasypus](/source/Prodasypus)''
** Genus [†](/source/Extinct)''[Proeutatus](/source/Proeutatus)''
** Genus [†](/source/Extinct)''[Prostegotherium](/source/Prostegotherium)''
** Genus [†](/source/Extinct)''[Pucatherium](/source/Pucatherium)''
** Genus [†](/source/Extinct)''[Punatherium](/source/Punatherium)''
** Genus [†](/source/Extinct)''[Stegotherium](/source/Stegotherium)''
** Genus [†](/source/Extinct)''[Stenotatus](/source/Stenotatus)''
** Genus [†](/source/Extinct)''[Utaetus](/source/Utaetus)''
** Genus [†](/source/Extinct)''[Vetelia](/source/Vetelia)''
** Subfamily [Dasypodinae](/source/Dasypodinae)
*** Genus [†](/source/Extinct)''[Anadasypus](/source/Anadasypus)''
*** Genus ''[Dasypus](/source/Dasypus)'' 
*** Genus [†](/source/Extinct)''[Nanoastegotherium](/source/Nanoastegotherium)''
*** Genus [†](/source/Extinct)''[Parastegosimpsonia](/source/Parastegosimpsonia)''
*** Genus [†](/source/Extinct)''[Pliodasypus](/source/Pliodasypus)''
*** Genus [†](/source/Extinct)''[Propraopus](/source/Propraopus)''
*** Genus [†](/source/Extinct)''[Riostegotherium](/source/Riostegotherium)''
*** Genus [†](/source/Extinct)''[Stegosimpsonia](/source/Stegosimpsonia)''
* '''Family [Chlamyphoridae](/source/Chlamyphoridae)''': glyptodonts and other armadillos
** Subfamily [Chlamyphorinae](/source/Chlamyphorinae): fairy armadillos
*** Genus ''[Calyptophractus](/source/Calyptophractus)''
*** Genus ''[Chlamyphorus](/source/Chlamyphorus)''
** Subfamily [Euphractinae](/source/Euphractinae): hairy, six-banded and pichi armadillos
*** Genus ''[Chaetophractus](/source/Chaetophractus)''
*** Genus [†](/source/Extinct)''[Doellotatus](/source/Doellotatus)''
*** Genus ''[Euphractus](/source/Euphractus)''
*** Genus [†](/source/Extinct)''[Macroeuphractus](/source/Macroeuphractus)''
*** Genus [†](/source/Extinct)''[Proeuphractus](/source/Proeuphractus)''
*** Genus [†](/source/Extinct)''[Paleuphractus](/source/Paleuphractus)''
*** Genus ''[Zaedyus](/source/Zaedyus)''
** Subfamily  [†](/source/Extinct)[Glyptodontinae](/source/Glyptodontinae): glyptodonts
*** Genus [†](/source/Extinct)''[Doedicurus](/source/Doedicurus)''
*** Genus [†](/source/Extinct)''[Glyptodon](/source/Glyptodon)''
*** Genus [†](/source/Extinct)''[Glyptotherium](/source/Glyptotherium)''
*** Genus [†](/source/Extinct)''[Hoplophorus](/source/Hoplophorus)''
*** Genus [†](/source/Extinct)''[Panochthus](/source/Panochthus)''
*** Genus [†](/source/Extinct)''[Parapropalaehoplophorus](/source/Parapropalaehoplophorus)''
*** Genus [†](/source/Extinct)''[Plaxhaplous](/source/Plaxhaplous)''
** Subfamily [Tolypeutinae](/source/Tolypeutinae): giant, three-banded and naked-tailed armadillos
*** Genus ''[Cabassous](/source/Cabassous)''
*** Genus [†](/source/Extinct)''[Kuntinaru](/source/Kuntinaru)''<ref name=Billetetal>{{Cite journal|author1=Guillaume Billet |author2=Lionel Hautier |author3=Christian de Muizon |author4=Xavier Valentin |year=2011 |title=Oldest cingulate skulls provide congruence between morphological and molecular scenarios of armadillo evolution |journal=Proceedings of the Royal Society |volume=278 |issue= 1719|pages= 2791–2797|doi=10.1098/rspb.2010.2443 |pmid=21288952 |pmc=3145180}}</ref>
*** Genus ''[Priodontes](/source/Priodontes)''
*** Genus ''[Tolypeutes](/source/Tolypeutes)''

{{clear}}
{{cladogram
|title=Cladogram of Cingulata<ref name=Delsuc2016/><ref name=Upham2019>
{{cite journal
 |first1=Nathan S. |last1=Upham
 |first2=Jacob A.  |last2=Esselstyn
 |first3=Walter    |last3=Jetz
 |year=2019
 |title=Inferring the mammal tree: Species-level sets of phylogenies for questions in ecology, evolution and conservation
 |journal=PLOS Biol
 |volume=17 |issue=12 |article-number=e3000494
 |pmid=31800571 |pmc=6892540
 |doi=10.1371/journal.pbio.3000494 |doi-access=free
}}
</ref><ref>
{{cite journal
 |last1=Gibb    |first1=Gillian C.      |last2=Condamine |first2=Fabien L.
 |last3=Kuch    |first3=Melanie         |last4=Enk       |first4=Jacob
 |last5=Moraes-Barros |first5=Nadia     |last6=Superina  |first6=Mariella
 |last7=Poinar  |first7=Hendrik N.      |last8=Delsuc    |first8=Frédéric
 |display-authors=6
 |year=2015
 |title=Shotgun mitogenomics provides a reference phylogenetic framework and timescale for living xenarthrans
 |journal=Molecular Biology and Evolution
 |volume=33 |issue=3 |pages=621–642
 |doi=10.1093/molbev/msv250
 |pmid=26556496 |pmc=4760074
}}
</ref>
|align=left
|width=520
|{{clade
|style=width:520px;font-size:90%;line-height:100%;
|label1= '''Cingulata''' 
|1={{clade
   |label1=[Dasypodidae](/source/Dasypodidae)
   |1={{clade
      |label1=''[Dasypus](/source/Dasypus)''
      |sublabel1=(¿[sensu lato](/source/sensu_lato)?)
      |1={{clade
         |label1=''D.'' (''[Hyperoambon](/source/Hyperoambon)'')
         |1={{clade
            |1=''[D. kappleri](/source/Dasypus_kappleri)''
            }}
         |label2=''D.'' (''[Dasypus](/source/Dasypus)'')
         |sublabel2=(¿[sensu stricto](/source/sensu_stricto)?)
         |2={{clade
            |label1=
            |1={{clade
               |label1=
               |1=''[D. septemcinctus](/source/Dasypus_septemcinctus)''
               |2=''[D. hybridus](/source/Dasypus_hybridus)''
               }}
            |2={{clade
               |label1=
               |1={{clade
                  |label1=
                  |1=''[D. yepesi](/source/Dasypus_yepesi)''
                  |2=''[D. sabanicola](/source/Dasypus_sabanicola)''
                  }}
               |2={{clade
                  |label1=
                  |1=''[D. novemcinctus](/source/Dasypus_novemcinctus)''
                  |2=''[D. pilosus](/source/Dasypus_pilosus)''
                  }}
               }}
            }}
         }}
     }}
  |label2=[Chlamyphoridae](/source/Chlamyphoridae)
  |2={{clade
     |label1=
     |1={{clade
        |label1=[Euphractinae](/source/Euphractinae)
        |1={{clade
           |label1=
           |1={{clade
              |label1= ''[Euphractus](/source/Euphractus)'' 
              |1=''[E. sexcinctus](/source/Euphractus_sexcinctus)''
              }}
           |2={{clade
              |label1=''[Zaedyus](/source/Zaedyus)''
              |1=''[Z. pichiy](/source/Zaedyus_pichiy)''
              |label2=''[Chaetophractus](/source/Chaetophractus)''
              |2={{clade
                 |label1=
                 |1=''[C. villosus](/source/Chaetophractus_villosus)''
                 |2={{clade
                    |label1=
                    |1=''[C. nationi](/source/Chaetophractus_nationi)''
                    |2=''[C. vellerosus](/source/Chaetophractus_vellerosus)''
                    }}
                 }}
              }}
           }}
        }}
     |2={{clade
        |label1=
        |1={{clade
           |label1=
           |1={{clade
              |label1={{extinct}}[Glyptodontinae](/source/Glyptodontinae)
              |1={{clade
                 |1={{clade
                    |label1= {{extinct}}''[Doedicurus](/source/Doedicurus)'' 
                    |1={{extinct}}''[D. clavicaudatus](/source/Doedicurus_clavicaudatus)''
                    }}
                 |2={{extinct}}(31 other extinct genera)
                 }}
              }}

        |2={{clade
           |label1=[Chlamyphorinae](/source/Chlamyphorinae)
           |1={{clade
             |label1=
             |1={{clade
                 |label1=''[Chlamyphorus](/source/Chlamyphorus)''
                 |1=''[C. truncatus](/source/Chlamyphorus_truncatus)''
                 }}
              |2={{clade
                 |label1=''[Calyptophractus](/source/Calyptophractus)''
                 |1=''[C. retusus](/source/Calyptophractus_retusus)''
                 }}
              }}
           |label2=[Tolypeutinae](/source/Tolypeutinae)
           |2={{clade
              |label1=
              |1={{clade
                 |label1=''[Priodontes](/source/Priodontes)''
                 |1=''[P. maximus](/source/Priodontes_maximus)''
                 }}
              |2={{clade
                 |label1=''[Tolypeutes](/source/Tolypeutes)''
                 |1={{clade
                    |label1=
                    |1=''[T. tricinctus](/source/Tolypeutes_tricinctus)''
                    |2=''[T. matacus](/source/Tolypeutes_matacus)''
                    }}
                 |label2=''[Cabassous](/source/Cabassous)''
                 |2={{clade
                    |label1=
                    |1=''[C. tatouay](/source/Cabassous_tatouay)''
                    |2={{clade
                       |label1=
                       |1=''[C. chacoensis](/source/Cabassous_chacoensis)''
                       |2={{clade
                          |1=''[C. centralis](/source/Cabassous_centralis)''
                          |2=''[C. unicinctus](/source/Cabassous_unicinctus)''
                       }}
                    }}
                 }}
              }}
           }}
         }}
       }}
     }}
   }}
 }}
}}
}}
{{clear}}

==References==
{{Reflist|2}}

{{Wikiquote}}
{{Wikispecies}}
{{Commons}}

{{mammals}}
{{Cingulata}}
{{Taxonbar|from=Q1242326}}

Category:Cingulata
Category:Mammal orders
Category:Taxa named by Johann Karl Wilhelm Illiger
Category:Extant Thanetian first appearances

---
Adapted from the Wikipedia article [Cingulata](https://en.wikipedia.org/wiki/Cingulata) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Cingulata?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.
