# Chabardella > Mediated Wiki article. Canonical URL: https://mediated.wiki/source/Chabardella > Markdown URL: https://mediated.wiki/source/Chabardella.md > Source: https://en.wikipedia.org/wiki/Chabardella > Source revision: 1332960546 > License: Creative Commons Attribution-ShareAlike 4.0 International (https://creativecommons.org/licenses/by-sa/4.0/) {{Short description|Extinct genus of mantis shrimp}} {{Italic title}} {{Speciesbox | fossil_range = [Late Carboniferous](/source/Late_Carboniferous) ([Stephanian](/source/Stephanian_(stage))), {{Fossil range|301.5|300.5}} | image = | genus = Chabardella | parent_authority = Racheboeuf, Schram and Vidal, 2009 | species = spinosa | authority = Racheboeuf, Schram and Vidal, 2009 }} '''''Chabardella''''' is an extinct [genus](/source/genus) of [mantis shrimp](/source/mantis_shrimp) which lived during the [Late Carboniferous](/source/Late_Carboniferous) in [France](/source/France). It was named in 2009, with '''''C. spinosa''''' as the type and only species. ==Discovery and naming== The fossil material of ''Chabardella'' was discovered in the Saint-Louis opencast pit in [Montceau-les-Mines](/source/Montceau-les-Mines), France, which dates to the [Stephanian](/source/Stephanian_(stage)) B and C substages of the [Late Carboniferous](/source/Late_Carboniferous) period. A research program on the fossil content of this site began in 2000, during which the only known specimen was found in a [concretion](/source/concretion). This fossil belongs to the [National Museum of Natural History, France](/source/National_Museum_of_Natural_History%2C_France) and was deposited in an off-site facility in [Autun](/source/Autun).{{Cite journal |last1=Racheboeuf |first1=Patrick R. |last2=Schram |first2=Frederick R. |last3=Vidal |first3=Muriel |date=2009 |title=New Malacostracan Crustacea from the Carboniferous (Stephanian) Lagerstätte of Montceau-Les-Mines, France |url=https://www.jstor.org/stable/29739132 |journal=Journal of Paleontology |volume=83 |issue=4 |pages=624–629 |doi=10.1666/08-171R.1 |jstor=29739132 |bibcode=2009JPal...83..624R |s2cid=130784496 |issn=0022-3360}} In 2009, Racheboeuf, Schram and Vidal described ''Chabardella spinosa'' as a new genus and species of [mantis shrimp](/source/mantis_shrimp) based on this fossil. The [holotype](/source/holotype) specimen, MNHN-SOT 28610, consists of the external mold of a weathered, near-complete [exoskeleton](/source/exoskeleton), preserving its left side. The [generic name](/source/Generic_name_(biology)) honors D. Chabard, who was curator of the Muséum d'histoire naturelle d'Autun and therefore in charge of the Montceau-les-Mines fossil collection. The [specific name](/source/Specific_name_(zoology)), ''spinosa'', references the rows of spines on the margins of the animal's segments. ==Description== ''Chabardella'' was a small [crustacean](/source/crustacean), with an arched body reaching a total length of around {{convert|30|mm|in|abbr=on}}. The animal's body was laterally compressed, being flattened from side to side and appearing narrow when viewed from above. Like other [mantis shrimp](/source/mantis_shrimp)s, ''Chabardella'' has [raptorial](/source/raptorial) maxillipeds (feeding appendages): the second to fifth segments of the [thorax](/source/Thorax_(arthropod_anatomy)) each bear a pair of enlarged [thoracopod](/source/thoracopod)s which would have been used to capture prey. The raptorial appendages of the frontmost pair are each {{convert|1.5|mm|in|abbr=on}} wide at the base, larger than those of the other three pairs (which are each {{convert|1|mm|in|abbr=on}} wide at the base). Though the [carapace](/source/carapace) is poorly preserved, it has been estimated to be {{convert|7.5|mm|in|abbr=on}} long and {{convert|5.5|mm|in|abbr=on}} high, and is known to cover the head and front part of the thorax, whereas the last four thoracic segments were left exposed. The segments of the thorax are short but tall, each measuring around {{convert|1.5|mm|in|abbr=on}} long and {{convert|4|mm|in|abbr=on}} tall. The [abdomen](/source/abdomen) is split into six segments, which decrease in size along the animal's length: the first abdominal segment is {{convert|2.5|mm|in|abbr=on}} long and {{convert|4|mm|in|abbr=on}} tall, whereas the last is only {{convert|1.3|mm|in|abbr=on}} long and {{convert|2|mm|in|abbr=on}} tall. The [telson](/source/telson) is elongated and subtriangular in shape, and the [uropod](/source/uropod) has a short, blade-like base. The rest of the tail fan is not preserved and thus poorly known. The hind margin of each thoracic and abdominal segment is almost vertical, and possesses ridges with five or six short, backward-pointing spines on each side. These spines are largest near the middle of the crustacean's back, decreasing in size down its sides. The hind segments also have slightly larger spines than the segments further front. These spines are a feature unique to ''Chabardella'', and are unknown in other [Paleozoic](/source/Paleozoic) mantis shrimps. ==Classification== Because the frontmost pair of [raptorial](/source/raptorial) appendages is larger than the latter pairs, Racheboeuf, Schram and Vidal (2009) recognized ''Chabardella'' as a close relative of ''[Gorgonophontes](/source/Gorgonophontes)'', which also has this feature. The authors thus placed ''Chabardella'' into [Gorgonophontidae](/source/Gorgonophontidae), making it the second genus assigned to this family. This placement was first questioned by Haug ''et al.'' (2010), which found ''Chabardella'', ''Gorgonophontes'' and the ''[Tyrannophontes](/source/Tyrannophontes)''+[Unipeltata](/source/Unipeltata) clade to form an unresolved [polytomy](/source/polytomy). However, they acknowledge this result may have been due to the lack of data on the tail fan of ''Chabardella'', which is poorly preserved.{{Cite journal |last1=Haug |first1=Joachim T. |last2=Haug |first2=Carolin |last3=Maas |first3=Andreas |last4=Kutschera |first4=Verena |last5=Waloszek |first5=Dieter |date=2010-09-21 |title=Evolution of mantis shrimps (Stomatopoda, Malacostraca) in the light of new Mesozoic fossils |journal=BMC Evolutionary Biology |language=en |volume=10 |issue=1 |pages=290 |doi=10.1186/1471-2148-10-290 |doi-access=free |issn=1471-2148 |pmc=2955030 |pmid=20858249|bibcode=2010BMCEE..10..290H }} Smith ''et al.'' (2023) conducted a [phylogenetic analysis](/source/phylogenetic_analysis) which also placed ''Chabardella'' in an unresolved polytomy, and further suggests the incomplete knowledge of ''[Daidal](/source/Daidal) pattoni'' may have also led to this result. The results of their [phylogenetic analyses](/source/phylogenetic_analyses) are displayed in the [cladogram](/source/cladogram) below:{{Cite journal |last1=Smith |first1=C.P.A. |last2=Aubier |first2=P. |last3=Charbonnier |first3=S. |last4=Laville |first4=T. |last5=Olivier |first5=N. |last6=Escarguel |first6=G. |last7=Jenks |first7=J.F. |last8=Bylund |first8=K.G. |last9=Fara |first9=E. |last10=Brayard |first10=A. |date=2023-03-31 |title=Closing a major gap in mantis shrimp evolution - first fossils of Stomatopoda from the Triassic |url=https://www.researchgate.net/publication/369959650 |journal=Bulletin of Geosciences |language=en |pages=95–110 |doi=10.3140/bull.geosci.1864 |issn=1802-8225|doi-access=free }} {{Cladogram of fossil mantis shrimp genera}} ==References== {{reflist}} {{taxonbar|from1=Q124387571|from2=Q124387655}} Category:Pennsylvanian animals of Europe Category:Stomatopoda Category:Fossil taxa described in 2009 Category:Prehistoric mantis shrimps Category:Carboniferous arthropods of Europe Category:Carboniferous crustaceans Category:Monotypic prehistoric crustacean genera --- Adapted from the Wikipedia article [Chabardella](https://en.wikipedia.org/wiki/Chabardella) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Chabardella?action=history)). 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