# Bathelium

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***Bathelium*** is a [genus](/source/Genus) of [lichen](/source/Lichen)-forming [fungi](/source/Fungi) in the family [Trypetheliaceae](/source/Trypetheliaceae).[1]

## Taxonomy

The genus was [circumscribed](/source/Circumscription_(taxonomy)) in 1803 by Swedish lichenologist [Erik Acharius](/source/Erik_Acharius), with *[Bathelium mastoideum](/source/Bathelium_mastoideum)* assigned as the [type species](/source/Type_species).[2]

## Description

Genus *Bathelium* features a thallus (a protective outer layer) usually with olive-green to brownish or greyish hues. This genus is distinguished by the form of its [ascomata](/source/Ascomata), the spore-producing structures, which possess apical [ostioles](/source/Ostiole) (openings at the top). These ascomata can be found either singly or in groups, and often form , a stroma made of both thallus tissue and bits of host tissue. They are noticeable, ranging from to , and show a brown-black colouration, though in rare instances, they may have a whitish (powdery) appearance. This is complemented by a peripheral layer made up of tightly packed cells.[3]

Within the ascomata, the , a tissue layer containing filamentous structures, is clear and [hyaline](/source/Hyaline) (translucent). It comprises thin, interconnecting filaments known as . The number from one to eight per [ascus](/source/Ascus). They are transversely [septate](/source/Septum) to (divided into compartments by cross walls), with thin septa and more or less angular . These spores are , meaning they have a spindle-like shape, with either acute (pointed) or rounded ends, and are not constricted at the median septum. They appear hyaline, do not react to [iodine](/source/Iodine) (IKI−), and are encased in a gelatinous sheath.[3]

No [conidiomata](/source/Conidiomata) (structures responsible for asexual reproduction) have been identified in this genus. In terms of chemical properties, *Bathelium* frequently contains [anthraquinones](/source/Anthraquinone), compounds often present internally in pseudostromata. [Lichexanthone](/source/Lichexanthone), a fluorescent [xanthone](/source/Xanthone) compound, is occasionally found within the genus.[3]

## Species

- *[Bathelium albidoporum](/source/Bathelium_albidoporum)* (Makhija & Patw.) R.C.Harris (1995)
- *[Bathelium austroafricanum](/source/Bathelium_austroafricanum)* (Zahlbr.) Aptroot & Lücking (2016)
- *[Bathelium boliviense](/source/Bathelium_boliviense)* Flakus & Aptroot (2016)[4] – Bolivia
- *[Bathelium carolinianum](/source/Bathelium_carolinianum)* (Tuck.) R.C.Harris (1995)
- *[Bathelium compositum](/source/Bathelium_compositum)* (Vain.) C.W.Dodge (1953)
- *[Bathelium connivens](/source/Bathelium_connivens)* (Nyl.) Trevis. (1861)
- *[Bathelium duplex](/source/Bathelium_duplex)* (Fée) C.W.Dodge (1953)
- *[Bathelium flavostiolatum](/source/Bathelium_flavostiolatum)* Flakus & Aptroot (2016)[4] – Bolivia
- *[Bathelium inspersomastoideum](/source/Bathelium_inspersomastoideum)* Flakus & Aptroot (2016)[4] – Bolivia
- *[Bathelium lineare](/source/Bathelium_lineare)* (C.W.Dodge) R.C.Harris (1995)
- *[Bathelium madreporiforme](/source/Bathelium_madreporiforme)* (Eschw.) Trevis. (1852)
- *[Bathelium mastoideum](/source/Bathelium_mastoideum)* Afzel. ex Ach. (1803)
- *[Bathelium megaleium](/source/Bathelium_megaleium)* Kremp. (1880)
- *[Bathelium meristosporum](/source/Bathelium_meristosporum)* Mont. & Bosch (1861)
- *[Bathelium mirabile](/source/Bathelium_mirabile)* Flakus, Kukwa & Aptroot (2016)[4] – Bolivia
- *[Bathelium nigroporum](/source/Bathelium_nigroporum)* (Makhija & Patw.) Aptroot & Lücking (2016)
- *[Bathelium oligosporum](/source/Bathelium_oligosporum)* (Mont. & Bosch) Trevis. (1861)
- *[Bathelium ostendatum](/source/Bathelium_ostendatum)* (Kremp.) Müll.Arg. (1880)
- *[Bathelium phaeomelodes](/source/Bathelium_phaeomelodes)* Müll.Arg. (1885)
- *[Bathelium porinosporum](/source/Bathelium_porinosporum)* Lücking, M.P.Nelsen & Gueidan (2016)[5]
- *[Bathelium pruinolucens](/source/Bathelium_pruinolucens)* Aptroot & Lücking (2016)
- *[Bathelium pruinosum](/source/Bathelium_pruinosum)* Flakus, Kukwa & Aptroot (2016)[4] – Bolivia
- *[Bathelium sphaericum](/source/Bathelium_sphaericum)* (C.W.Dodge) R.C.Harris (1995)
- *[Bathelium subalbens](/source/Bathelium_subalbens)* (Nyl.) C.W.Dodge (1953)
- *[Bathelium sundaicum](/source/Bathelium_sundaicum)* Müll.Arg. (1891)
- *[Bathelium tuberculosum](/source/Bathelium_tuberculosum)* (Makhija & Patw.) R.C.Harris (1995)
- *[Bathelium varium](/source/Bathelium_varium)* (Fée) Trevis. (1861)
- *[Bathelium velatum](/source/Bathelium_velatum)* Müll.Arg. (1882)

## References

1. Wijayawardene, Nalin; Hyde, Kevin; Al-Ani, Laith Khalil Tawfeeq; Somayeh, Dolatabadi; Stadler, Marc; Haelewaters, Danny et al. (2020). "Outline of Fungi and fungus-like taxa". *Mycosphere*. **11**: 1060–1456. [doi:10.5943/mycosphere/11/1/8](https://doi.org/10.5943/mycosphere/11/1/8). [hdl:10481/61998](https://hdl.handle.net/10481/61998)

1. Acharius, E. (1803). [*Methodus qua Omnes Detectos Lichenes Secundum Organa Carpomorpha ad Genera, Species et Varietates Redigere atque Observationibus Illustrare Tentavit Erik Acharius*](https://www.biodiversitylibrary.org/page/44218201) (in Latin). Stockholm: impensis F.D.D. Ulrich. p. 111.

1. Aptroot, André & Lücking, Robert (2016). "A revisionary synopsis of the Trypetheliaceae (Ascomycota: Trypetheliales)". *The Lichenologist*. **48** (6): 763–982. [doi:10.1017/s0024282916000487](https://doi.org/10.1017/s0024282916000487)

1. Flakus, Adam; Kukwa, Martin; Aptroot, André (2016). "Trypetheliaceae of Bolivia: an updated checklist with descriptions of twenty-four new species". *The Lichenologist*. **48** (6): 661–692. [doi:10.1017/S0024282915000559](https://doi.org/10.1017/S0024282915000559)

1. Lücking, Robert; Nelsen, Matthew P.; Aptroot, André; Benatti, Michel Navarro; Binh, Nguyen Quoc; Gueidan, Cecile; Gutiérrez, Martha Cecilia; Jungbluth, Patricia; Lumbsch, H. Thorsten; Marcelli, Marcelo P.; Moncada, Bibiana; Naksuwankul, Khwanruan; Orozco, Thelma; Salazar-Allen, Noris; Upreti, Dalip K. (2016). "A pot-pourri of new species of Trypetheliaceae resulting from molecular phylogenetic studies". *The Lichenologist*. **48** (6): 639–660. [doi:10.1017/S0024282916000475](https://doi.org/10.1017/S0024282916000475)

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Adapted from the Wikipedia article [Bathelium](https://en.wikipedia.org/wiki/Bathelium) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Bathelium?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.
