# Asplenium anceps

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***Asplenium anceps*** is a diploid [fern](/source/Fern) of family [Aspleniaceae](/source/Aspleniaceae) and one of the ancestors of the ferns that form the trichomanes complex. It lives exclusively in the three northernmost archipelagoes of the [Macaronesian](/source/Macaronesia) region, that is, is an endemic macaronesian fern. Its fronds are leathery and plastic and [rachis](/source/Rachis) is very thick, bright reddish brown and is traversed throughout its length of three wings, two on the upper surface to draw a groove and a third on the lower surface which is characteristic and unique to this species, since all other species of the trichomanes complex without. A typical feature of this fern, which he shares with all its hybrid offspring (*Asplenium azoricum*, *Asplenium azomanes* and *Asplenium* X *tubalense*) is the existence of a small atrium on the basis of medium and less pinnae directed toward the apex of the blade with one or two sori on its underside.

## Taxonomy

A global phylogeny of *Asplenium* published in 2020 divided the genus into eleven clades,[1] which were given informal names pending further taxonomic study. *A. anceps* belongs to the "*A. trichomanes* subclade" of the "*A. trichomanes* clade".[2] The *A. trichomanes* clade has a worldwide distribution. Members of the clade grow on rocks and have once-pinnate leaf blades with slender, chestnut- to dark-brown stalks.[3] The *A. trichomanes* subclade also has brown, membraneous wings or projections on the side of the rachis. Relatives within the subclade include the (polyphyletic) *[A. trichomanes](/source/Asplenium_trichomanes)* and *[A. tripteropus](/source/Asplenium_tripteropus)*.[4]

## Habitat

It grows in the shade of laurel forests and high mountain pine forests between the stones of the walls and crevices of volcanic rock oriented north and northwest, on a base of moss and lichen.

## Distribution

It lives in [Azores](/source/Azores), [Madeira](/source/Madeira) and [Canary Islands](/source/Canary_Islands). Since 1972 there have been no records of this plant in the Azores.[5]

## Hybrids

- [*Asplenium azoricum*](/source/Asplenium_azoricum): allotetraploid hybrid by cross between *A. anceps* and an individual of trichomanes complex.

## References

1. Xu et al. 2020, p. 27.

1. Xu et al. 2020, p. 39.

1. Xu et al. 2020, p. 44.

1. Xu et al. 2020, p. 45.

1. ["Asplenium anceps Lowe ex Hook. & Grev."](https://acores.flora-on.pt/#/1Asplenium+anceps). [Flora-On](/source/Flora-On). Retrieved 4 March 2022.

## Sources

- Xu, Ke-Wang; Zhang, Liang; Rothfels, Carl J.; Smith, Alan R.; Viane, Ronald; Lorence, David; Wood, Kenneth R.; Chen, Cheng-Wei; Knapp, Ralf; Zhou, Lin; Lu, Ngan Thi; Zhou, Xin-Mao; Wei, Hong-Jin; Fan, Qiang; Chen, Su-Fang; Cicuzza, Daniele; Gao, Xin-Fen; Li, Wen-Bo; Zhang, Li-Bing (2020). ["A global plastid phylogeny of the fern genus Asplenium (Aspleniaceae)"](https://onlinelibrary.wiley.com/doi/abs/10.1111/cla.12384). *[Cladistics](/source/Cladistics_(journal))*. **36** (1): 22–71. [doi:10.1111/cla.12384](https://doi.org/10.1111/cla.12384). [PMID 34618950](https://pubmed.ncbi.nlm.nih.gov/34618950). [S2CID 201197385](https://api.semanticscholar.org/CorpusID:201197385)
- [Asplenium anceps in Azores Islands](http://www.azoresbioportal.angra.uac.pt/listagens.php?lang=es&start=121&end=160&sstr=4&id=F00011) [Archived](https://web.archive.org/web/20120304234655/http://www.azoresbioportal.angra.uac.pt/listagens.php?lang=es&start=121&end=160&sstr=4&id=F00011) 2012-03-04 at the Wayback Machine
- [Asplenium anceps in Red List of "Flora Vascular Española"](https://web.archive.org/web/20091214133957/http://www.mma.es/portal/secciones/biodiversidad/inventarios/inb/flora_vascular/pdf/ListaRojaFlora08.pdf)
- [See photos of Asplenium anceps](https://web.archive.org/web/20110719114843/http://jardin-mundani.info/endemismos-maderenses/flora-maderense-macaronesica.htm)
- [More photos of Asplenium anceps](https://web.archive.org/web/20110719114832/http://jardin-mundani.info/falgueres/aspleniaceas/asplenium.htm)

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Adapted from the Wikipedia article [Asplenium anceps](https://en.wikipedia.org/wiki/Asplenium_anceps) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Asplenium_anceps?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.