# Ark clam

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Family of molluscs

Ark clam Shell specimens of Anadara antiquata Scientific classification Kingdom: Animalia Phylum: Mollusca Class: Bivalvia Order: Arcida Superfamily: Arcoidea Family: Arcidae Lamarck, 1809 Genera See text

**Ark clam** is the [common name](/source/Common_name) for a family of small to large-sized saltwater clams or [marine](/source/Marine_(ocean)) [bivalve](/source/Bivalve) [molluscs](/source/Mollusc) in the [family](/source/Family_(taxonomy)) **Arcidae**. Generally less than 80 mm long, ark clams vary both in shape and size. They number about 200 species worldwide.[1]

The group is known as "ark shells" because species such as *[Arca](/source/Arca_(bivalve))* have a large flat area between the [umbones](/source/Umbo_(bivalve)) which, in an undamaged shell, somewhat resembles a [deck](/source/Deck_(ship)), with the rest of the shell perhaps illustrating an ancient wooden boat such as [Noah's Ark](/source/Noah's_Ark) is thought to have been.

The thick, ridged shells of ark clams are often white, cream or tan,[1] but in some species, the shell is striped with, tinted with, or completely colored, a rich brown. In life the shell of most species has a top shell layer that is thick brown [periostracum](/source/Periostracum) affixed to the harder calcareous part of the shell. In some species such as *[Barbatia](/source/Barbatia)*, this outer horny covering is tufted at the end of the shell into something that resembles a beard, hence the name *Barbatia* or bearded one. The thick outer skin or [periostracum](/source/Periostracum) of an ark clam can act as [camouflage](/source/Camouflage), such that the shells can sometimes look like stones when lying on the bottom.

One fossil [valve](/source/Valve_(mollusc)) of *[Anadara](/source/Anadara)* from [Cyprus](/source/Cyprus), dating to the [Pliocene](/source/Pliocene) [Epoch](/source/Epoch_(geology)) of the [geologic timescale](/source/Geologic_timescale), approx. 5.3-2.5 million years [BP](/source/Before_Present)

All ark shells have a long straight hinge line with a single row of numerous small and unspecialized "teeth". This is known as a "taxodont dentition" and represents an ancient ancestor. This kind of hinge line is also found in the bivalve families [Glycymerididae](/source/Glycymerididae), [Nuculidae](/source/Nuculidae) and [Nuculanidae](/source/Nuculanidae).

Ark clams are distinct from other clams in having red blood pigments ([hemoglobin](/source/Hemoglobin)) that facilitate the transport of [oxygen](/source/Oxygen) to their tissues and enable them to inhabit more [hypoxic environments](/source/Hypoxia_(environmental)).[1]

Ark clams reach reproductive maturity when they are about twelve months old and about 20 mm long. The [spawning](/source/Spawn_(biology)) cycle typically begins in the rainy season. Ark Clams are *broadcast spawners*, that is, eggs and sperm are released into open water where fertilization occurs. The fertilized eggs develop rapidly into [planktonic](/source/Planktonic) [larvae](/source/Larva) that drift with ocean currents for eight to ten days during which 99.9% of the larvae are consumed or perish. Eventually the survivors settle to suitable sites of the sea floor where they develop into juvenile clams. Only one percent of these juveniles will survive to become a mature adult. Ark Clams have a maximum life span of about six years.[1]

## Human use

Japanese *akagai* (*Anadara broughtonii*) served as [sushi](/source/Sushi)

Boiled ark clams served in [Tanjong Pagar](/source/Tanjong_Pagar), [Singapore](/source/Singapore)

Ark clams are edible mollusks that have been consumed since pre-historic times and are still consumed today. Numerous recipes for ark clams have been published. Recreational foragers should follow seafood safety guidelines and shellfish harvesting restrictions, and should be aware of any [harmful algal blooms](/source/Harmful_algal_bloom) that may contaminate shellfish with biotoxins. Shellfish collected in urban areas should not be eaten raw.

*[Tegillarca granosa](/source/Tegillarca_granosa)* was used as a food by Indigenous peoples living on the northern Australian coastline through at least the past ~4500 years, with extensive evidence preserved in the form of [shell mound](/source/Shell_mound) sites.[2] Large ark clams, such as *[Arca zebra](/source/Arca_zebra)*, are commonly used as bait, as well as food, throughout the [Caribbean](/source/Caribbean). In Japan, red Ark clams, called Akagai, are used in [sushi](/source/Sushi) or [sashimi](/source/Sashimi).[3] Some ark clams species, such as the [blood cockle](/source/Blood_cockle) (*[Anadara granosa](/source/Anadara_granosa)*, a.k.a. *[Tegillarca granosa](/source/Tegillarca_granosa)*) are raised in [aquaculture](/source/Aquaculture), e.g. in the estuaries of China's [Fujian](/source/Fujian) coast.[4] In the U.S. limited quantities of wild ark clams have been harvested in [North Carolina](/source/North_Carolina) and [Virginia](/source/Virginia) for ethnic markets and aquaculture has been explored. [5]

In the south Pacific region, *Ark* clams are still gathered by indigenous people as an important subsistence food. Ark clams are harvested, mostly by women, by **[gleaning](/source/Gleaning)** intertidal zones.[1] To maintain the Ark Clam fishery, several communities in [Fiji](/source/Fiji) are imposing a minimum size limit of 3 cm, closures during spawning periods, and establishing "no-take" areas.[1]

## Genera

Numerous valves of arcids, genus *[Senilia](/source/Senilia)*, washed up on the beach in [Senegal](/source/Senegal)

Genera within the family Arcidae include:

- *[Acar](/source/Acar_(bivalve))* [Gray](/source/John_Edward_Gray), 1857

- *[Anadara](/source/Anadara)* [Gray](/source/John_Edward_Gray), 1847

- *[Arca](/source/Arca_(bivalve))* [Linnaeus](/source/Carl_Linnaeus), [1758](/source/10th_edition_of_Systema_Naturae)

- *[Barbatia](/source/Barbatia)* [Gray](/source/John_Edward_Gray), 1847

- *[Bathyarca](/source/Bathyarca)* Kobelt, 1891

- *[Bentharca](/source/Bentharca)* Verrill and Bush, 1898

- *[Larkinia](/source/Larkinia)* Reinhart, 1935

- *[Samacar](/source/Samacar)* [Iredale](/source/Tom_Iredale), 1936

- *[Senilia](/source/Senilia)* [Linnaeus](/source/Carl_Linnaeus), [1758](/source/10th_edition_of_Systema_Naturae)

- *[Tegillarca](/source/Tegillarca)* [Iredale](/source/Tom_Iredale), 1939

- *[Palestinarca](/source/Palestinarca)*†

## References

1. ^ [***a***](#cite_ref-:0_1-0) [***b***](#cite_ref-:0_1-1) [***c***](#cite_ref-:0_1-2) [***d***](#cite_ref-:0_1-3) [***e***](#cite_ref-:0_1-4) [***f***](#cite_ref-:0_1-5) The Pacific Community (SPC) (2021). ["Information Sheet for Fishing Communities #22: Ark Clams"](https://lmmanetwork.org/wp-content/uploads/2021/08/Anon_12_ISFC_22_Anadara.pdf) (PDF). Retrieved June 2, 2024.

1. **[^](#cite_ref-2)** Patrick, Faulkner (2013). *Life on the margins : an archaeological investigation of late Holocene economic variability, Blue Mud Bay, Northern Australia*. Acton, A.C.T. [ISBN](/source/ISBN_(identifier)) [9781925021103](https://en.wikipedia.org/wiki/Special:BookSources/9781925021103). [OCLC](/source/OCLC_(identifier)) [850906221](https://search.worldcat.org/oclc/850906221).{{[cite book](https://en.wikipedia.org/wiki/Template:Cite_book)}}: CS1 maint: location missing publisher ([link](https://en.wikipedia.org/wiki/Category:CS1_maint:_location_missing_publisher))

1. **[^](#cite_ref-3)** ["Akagai (Ark Shell Clam)"](https://www.sushifaq.com/sushi-sashimi-info/the-many-different-types-of-sushi-items-and-their-japanese-names/akagai-surf-clam/). *The Sushi FAQ*. Retrieved 2024-06-02.

1. **[^](#cite_ref-4)** Ruǎn Jīnshān; Li Xiùzhū; Lín Kèbīng; Luō Dōnglián; Zhōu Chén; Cài Qīnghǎi (阮金山;李秀珠;林克冰;罗冬莲;周宸;蔡清海), [安海湾南岸滩涂养殖贝类死亡原因调查分析](http://mall.cnki.net/magazine/Article/FJSC200504006.htm) [Archived](https://web.archive.org/web/20200717121301/https://mall.cnki.net/magazine/Article/FJSC200504006.htm) 2020-07-17 at the [Wayback Machine](/source/Wayback_Machine) (Analysis of the causes of death of farmed shellfish on the mudflats in the southern part of Anhai Bay), 《福建水产》 (*Fujian Aquaculture*), 2005-04

1. **[^](#cite_ref-5)** University of Florida's Institute of Food and Agricultural Sciences (UF/IFAS). ["Investigation of Blood Ark and Ponderous Ark Culture and Marketability"](https://shellfish.ifas.ufl.edu/projects/shellfish-aquaculture-production-and-management/blood-and-ponderous-ark/). *Florida Shellfish Aquaculture Online Resource Guide*. Retrieved 2024-06-02.

## External links

[Wikispecies](/source/Wikispecies) has information related to ***[Arcidae](https://species.wikimedia.org/wiki/Arcidae)***.

- [ITIS](https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=79326)

- [Archerd Shell Collection, Ark clams](http://shells.tricity.wsu.edu/ArcherdShellCollection/Bivalvia/Arcidae.html) [Archived](https://web.archive.org/web/20120606032131/http://shells.tricity.wsu.edu/ArcherdShellCollection/Bivalvia/Arcidae.html) 2012-06-06 at the [Wayback Machine](/source/Wayback_Machine)

Taxon identifiers Arcidae Wikidata: Q632636 Wikispecies: Arcidae ADW: Arcidae AFD: Arcidae BOLD: 696 CoL: 7NFZ3 EoL: 46465774 GBIF: 3483 iNaturalist: 85598 IRMNG: 105092 ITIS: 79326 NBN: NHMSYS0021055016 NCBI: 6553 NZOR: bc0f415d-09b7-4650-b871-eb7815f71152 Open Tree of Life: 499758 Paleobiology Database: 59188 WoRMS: 208

Authority control databases National United States Israel Other Yale LUX

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Adapted from the Wikipedia article [Ark clam](https://en.wikipedia.org/wiki/Ark_clam) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Ark_clam?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.