# Aphodius

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***Aphodius*** is a [genus](/source/Genus) of beetles in the family [Scarabaeidae](/source/Scarabaeidae). In most species both the adults and [larvae](/source/Larva) are [coprophagous](/source/Coprophagous) (dung feeding)[1] although some species have [herbivorous](/source/Herbivore) or [saprophagous](/source/Saprotrophic_nutrition) larvae.[2] *Aphodius* species typically dominate [dung beetle](/source/Dung_beetle) communities in north temperate ecosystems.[3] Most species are functionally classified as endocoprids, also known as dwellers, because the larvae live and feed within the dung pat itself.[4]

With a discovery of an earlier published version by Hellwig, the correct citation for this genus is now: *Aphodius* Hellwig, 1798 (p. 101; 24 January). Type species by subsequent designation by Latreille (1810): *Scarabaeus fimetarius* Linnaeus, 1758.[5]

## Species

These 44 species belong to the genus *Aphodius*, including 21 extinct species.[6]

- *[Aphodius beloni](/source/Aphodius_beloni)* Mulsant & Godart, 1879
- *[Aphodius calichromus](/source/Aphodius_calichromus)* Balthasar, 1932
- *[Aphodius cardinalis](/source/Aphodius_cardinalis)* Reitter, 1892
- *[Aphodius clypeatus](/source/Aphodius_clypeatus)* Fischer von Waldheim, 1821
- *[Aphodius coniugatus](/source/Aphodius_coniugatus)* (Panzer, 1795)
- *[Aphodius corallifer](/source/Aphodius_corallifer)* Koshantschikov, 1913
- *[Aphodius crux](/source/Aphodius_crux)* Wiedemann, 1823
- *[Aphodius elegans](/source/Aphodius_elegans)* Allibert, 1847
- *[Aphodius fasciger](/source/Aphodius_fasciger)* Harold, 1881
- *[Aphodius fimetarius](/source/Aphodius_fimetarius)* (Linnaeus, 1758)
- *[Aphodius foetidus](/source/Aphodius_foetidus)* (Herbst, 1783)
- *[Aphodius guangdongensis](/source/Aphodius_guangdongensis)* Maté, 2008
- *[Aphodius irregularis](/source/Aphodius_irregularis)* Westwood, 1839
- *[Aphodius marginatus](/source/Aphodius_marginatus)* Fischer von Waldheim, 1842
- *[Aphodius micros](/source/Aphodius_micros)* Walker, 1871
- *[Aphodius minatorius](/source/Aphodius_minatorius)* Péringuey, 1908
- *[Aphodius pedellus](/source/Aphodius_pedellus)* (Degeer, 1774)
- *[Aphodius pedrosi](/source/Aphodius_pedrosi)* Wollaston, 1854
- *[Aphodius plasoni](/source/Aphodius_plasoni)* Käufel, 1914
- *[Aphodius reginae](/source/Aphodius_reginae)* Král, 1997
- *[Aphodius spadix](/source/Aphodius_spadix)* Schmidt, 1916
- *[Aphodius swaneticus](/source/Aphodius_swaneticus)* Reitter, 1892
- *[Aphodius thoracicus](/source/Aphodius_thoracicus)* Fischer von Waldheim, 1842
- † *[Aphodius aboriginalis](/source/Aphodius_aboriginalis)* Wickham, 1912
- † *[Aphodius anteactus](/source/Aphodius_anteactus)* Krell, 2000
- † *[Aphodius bosniaskii](/source/Aphodius_bosniaskii)* Handlirsch, 1907
- † *[Aphodius brevipennis](/source/Aphodius_brevipennis)* Heer, 1862
- † *[Aphodius charauxi](/source/Aphodius_charauxi)* Piton, 1940
- † *[Aphodius cretaceous](/source/Aphodius_cretaceous)* Nikolajev, 2008
- † *[Aphodius florissantensis](/source/Aphodius_florissantensis)* Wickham, 1911
- † *[Aphodius granarioides](/source/Aphodius_granarioides)* Wickham, 1913
- † *[Aphodius helvolus](/source/Aphodius_helvolus)* Statz, 1952
- † *[Aphodius inundatus](/source/Aphodius_inundatus)* Wickham, 1914
- † *[Aphodius krantzi](/source/Aphodius_krantzi)* Heyden & Heyden, 1866
- † *[Aphodius laminicola](/source/Aphodius_laminicola)* Wickham, 1910
- † *[Aphodius mediaevus](/source/Aphodius_mediaevus)* Wickham, 1914
- † *[Aphodius meyeri](/source/Aphodius_meyeri)* Heer, 1847
- † *[Aphodius praeemptor](/source/Aphodius_praeemptor)* Wickham, 1913
- † *[Aphodius precursor](/source/Aphodius_precursor)* Horn, 1876
- † *[Aphodius schlickumi](/source/Aphodius_schlickumi)* Statz, 1952
- † *[Aphodius senex](/source/Aphodius_senex)* Wickham, 1914
- † *[Aphodius shoshonis](/source/Aphodius_shoshonis)* Wickham, 1912
- † *[Aphodius theobaldi](/source/Aphodius_theobaldi)* Krell, 2000
- † *[Aphodius vectis](/source/Aphodius_vectis)* Krell, 2019

The species of the following genera were formerly classified in *Aphodius*:[7][8]

- *[Acrossus](/source/Acrossus)* Mulsant, 1842
- *[Agoliinus](/source/Agoliinus)* Schmidt, 1913
- *[Alloblackburneus](/source/Alloblackburneus)* Bordat, 2009
- *[Ballucus](/source/Ballucus)* Gordon & Skelley, 2007
- *[Blackburneus](/source/Blackburneus)* Schmidt, 1913
- *[Calamosternus](/source/Calamosternus)* Motschulsky, 1859
- *[Caligodorus](/source/Caligodorus)* Gordon & Skelley, 2007
- *[Cephalocyclus](/source/Cephalocyclus)* Dellacasa et al., 1998
- *[Chilothorax](/source/Chilothorax)* Motschulsky, 1859
- *[Cinacanthus](/source/Cinacanthus)* Schmidt, 1913
- *[Coelotrachelus](/source/Coelotrachelus)* Schmidt, 1913
- *[Colobopterus](/source/Colobopterus)* Mulsant, 1842
- *[Cryptoscatomaseter](/source/Cryptoscatomaseter)* Gordon & Skelley, 2007
- *[Dellacasiellus](/source/Dellacasiellus)* Gordon & Skelley, 2007
- *[Dialytodius](/source/Dialytodius)* Gordon & Skelley, 2007
- *[Drepanocanthoides](/source/Drepanocanthoides)* Schmidt, 1913
- *[Eupleurus](/source/Eupleurus)* Mulsant, 1842
- *[Flaviellus](/source/Flaviellus)* Gordon & Skelley, 2007
- *[Geomyphilus](/source/Geomyphilus)* Gordon & Skelley, 2007
- *[Haroldiellus](/source/Haroldiellus)* Gordon & Skelley, 2007
- *[Hornosus](/source/Hornosus)* Dellacasa, Dellacasa & Gordon, 2015
- *[Irrasinus](/source/Irrasinus)* Gordon & Skelley, 2007
- *[Labarrus](/source/Labarrus)* Mulsant & Rey, 1869
- *[Lechorodius](/source/Lechorodius)* Gordon & Skelley, 2007
- *[Liothorax](/source/Liothorax)* Motschulsky, 1859
- *[Luxolinus](/source/Luxolinus)* Gordon & Skelley, 2007
- *[Maculaphodius](/source/Maculaphodius)* Gordon & Skelley, 2007
- *[Melinopterus](/source/Melinopterus)* Mulsant, 1842
- *[Mendidius](/source/Mendidius)* Harold, 1868
- *[Merogyrus](/source/Merogyrus)* Gordon & Skelley, 2007
- *[Neotrichonotulus](/source/Neotrichonotulus)* Dellacasa et al., 2004
- *[Nialaphodius](/source/Nialaphodius)* Kolbe, 1908
- *[Orodaliscoides](/source/Orodaliscoides)* Schmidt, 1913
- *[Oscarinus](/source/Oscarinus)* Gordon & Skelley, 2007
- *[Otophorus](/source/Otophorus)* Mulsant, 1842
- *[Oxyomoides](/source/Oxyomoides)* Dellacasa et al., 2016
- *[Pardalosus](/source/Pardalosus)* Gordon & Skelley, 2007
- *[Phaeaphodius](/source/Phaeaphodius)* Reitter, 1892
- *[Planolinellus](/source/Planolinellus)* Dellacasa & Dellacasa, 2005
- *[Planolinoides](/source/Planolinoides)* Dellacasa & Dellacasa, 2005
- *[Planolinus](/source/Planolinus)* Mulsant & Rey, 1869
- *[Pseudagolius](/source/Pseudagolius)* Schmidt, 1913
- *[Rugaphodius](/source/Rugaphodius)* Gordon & Skelley, 2007
- *[Scabrostomus](/source/Scabrostomus)* Gordon & Skelley, 2007
- *[Schaefferellus](/source/Schaefferellus)* Gordon & Skelley, 2007
- *[Setodius](/source/Setodius)* Gordon & Skelley, 2007
- *[Stenotothorax](/source/Stenotothorax)* Schmidt, 1913
- *[Strigodius](/source/Strigodius)* Gordon & Skelley, 2007
- *[Tetraclipeoides](/source/Tetraclipeoides)* Schmidt, 1913
- *[Teuchestes](/source/Teuchestes)* Mulsant, 1842
- *[Trichaphodioides](/source/Trichaphodioides)* Paulian, 1942
- *[Trichonotulus](/source/Trichonotulus)* Bedel, 1911

## References

1. Valiela, Ivan (1974). "Composition, food webs, and population limitation in dung arthropod communities during invasion and succession". *American Midland Naturalist*. **92**: 370–385 [380]. [doi:10.2307/2424302](https://doi.org/10.2307/2424302). [JSTOR 2424302](https://www.jstor.org/stable/2424302)

1. Hanski and Cambefort (1991), p. 83.

1. Hanski and Cambefort (1991), p. 75.

1. Finn, J. A. & Gittings, T. (2003). "A review of competition in north temperate dung beetle communities". *Ecological Entomology*. **28** (1): 1–13. [doi:10.1046/j.1365-2311.2002.00487.x](https://doi.org/10.1046/j.1365-2311.2002.00487.x)

1. ALONSO-ZARAZAGA, M.A. & KRELL, F.T. 2011. Change of authorship of *Aphodius* and *Oryctes* to Hellwig, 1798 (Insecta: Coleoptera: Scarabaeidae). Zootaxa 3060: 67–68. [https://www.mapress.com/zt/article/view/zootaxa.3060.1.5/20827](https://www.mapress.com/zt/article/view/zootaxa.3060.1.5/20827)

1. ["*Aphodius* Hellwig, 1798"](https://www.catalogueoflife.org/data/taxon/YFV). *Catalogue of Life*. Retrieved 2021-12-25.

1. Gordon, Robert D. & Skelley, Paul E. (2007). "A monograph of the Aphodiini inhabiting the United States and Canada (Coleoptera: Scarabaeidae: Aphodiini)". *Memoirs of the American Entomological Institute*. **79**: 1–580. ISBN 978-1-887988-23-0.

1. ["*Aphodius* Genus information"](https://bugguide.net/node/view/41325). *Bugguide*. Retrieved 2021-12-25.

- *Dung Beetle Ecology*. Princeton University Press. 1991. [doi:10.1515/9781400862092](https://doi.org/10.1515/9781400862092). ISBN 9781400862092.

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Adapted from the Wikipedia article [Aphodius](https://en.wikipedia.org/wiki/Aphodius) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Aphodius?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.