# Amphipyrinae > Mediated Wiki article. Canonical URL: https://mediated.wiki/source/Amphipyrinae > Markdown URL: https://mediated.wiki/source/Amphipyrinae.md > Source: https://en.wikipedia.org/wiki/Amphipyrinae > Source revision: 1342307138 > License: Creative Commons Attribution-ShareAlike 4.0 International (https://creativecommons.org/licenses/by-sa/4.0/) Subfamily of moths Amphipyrinae Psaphida Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Superfamily: Noctuoidea Family: Noctuidae Subfamily: Amphipyrinae Guenée, 1837 Tribes Amphipyrini Guenée, 1837 Psaphidini Grote, 1896 *Amphipyra pyramidoides*, Copper underwing, Oklahoma *Copivaleria grotei*, Grote's Sallow, Ontario, Canada *Emarginea percara*, Beloved emarginea, Oklahoma **Amphipyrinae** is a subfamily of [owlet moths](/source/Owlet_moth) in the family [Noctuidae](/source/Noctuidae). There are more than 50 genera and 210 described species in Amphipyrinae, although the classifications are likely to change over time. This subfamily has been used as a catchall for members of Noctuidae that don't fit well into other subfamilies. As such, many of its members lack morphological traits that would allow assignment into one of the other subfamilies. Genetic analysis conducted on Amphipyrinae is improving the classification of these genera.[1] In 2021, phylogenetic research resulted in 11 genera being transferred from Amphipyrinae to six different subfamilies, and it is expected that more members of Amphipyrinae will be reassigned as further research is done.[2] Psaphidini is included here as a tribe, but is sometimes treated as the subfamily Psaphidinae. The Australian genera in Acronictinae are sometimes considered part of Amphipyrinae.[1][2] ## Genera These 52 genera belong to the subfamily Amphipyrinae:[3][4][5][2] **Tribe [Amphipyrini](https://en.wikipedia.org/w/index.php?title=Amphipyrini&action=edit&redlink=1) Guenée, 1837** - - *[Amphipyra](/source/Amphipyra)* Ochsenheimer, 1816 **Tribe [Psaphidini](/source/Psaphidini) Grote, 1896** - - Subtribe [Feraliina](https://en.wikipedia.org/w/index.php?title=Feraliina&action=edit&redlink=1) Poole, 1995 - *[Apsaphida](/source/Apsaphida)* Franclemont, 1973 - *[Feralia](/source/Feralia_(moth))* Grote, 1874 - *[Miracavira](/source/Miracavira)* Franclemont, 1937 - *[Paratrachea](/source/Paratrachea)* Hampson, 1908 - *[Viridemas](/source/Viridemas)* Smith, 1908 - Subtribe [Nocloina](https://en.wikipedia.org/w/index.php?title=Nocloina&action=edit&redlink=1) Poole, 1995 - *[Emarginea](/source/Emarginea)* Guenée, 1852 - *[Euamiana](/source/Euamiana)* Barnes & Benjamin, 1927 - *[Lythrodes](/source/Lythrodes)* Smith, 1903 - *[Nocloa](/source/Nocloa)* Smith, 1906 - *[Oslaria](/source/Oslaria)* Dyar, 1904 - *[Paramiana](/source/Paramiana)* Barnes & Benjamin, 1924 - *[Petalumaria](/source/Petalumaria)* Buckett & Bauer, 1968 - *[Prothrinax](/source/Prothrinax)* Hampson, 1908 - *[Redingtonia](/source/Redingtonia)* Barnes & McDunnough, 1912 - *[Ruacodes](/source/Ruacodes)* Hampson, 1908 - *[Walterella](/source/Walterella)* Dyar, 1921 - Subtribe [Psaphidina](https://en.wikipedia.org/w/index.php?title=Psaphidina&action=edit&redlink=1) Grote, 1896 - *[Brachionycha](/source/Brachionycha)* Hübner, 1819 - *[Copivaleria](/source/Copivaleria)* Grote, 1883 - *[Pleromella](/source/Pleromella)* Dyar, 1921 - *[Provia](/source/Provia_(moth))* Barnes & McDunnough, 1910 - *[Psaphida](/source/Psaphida)* Walker, 1865 - *[Pseudocopivaleria](/source/Pseudocopivaleria)* Buckett & Bauer, 1966 - Subtribe [Triocnemidina](/source/Triocnemidina) Poole, 1995 - *[Crimona](/source/Crimona)* Smith, 1902 - *[Oxycnemis](/source/Oxycnemis)* Grote, 1882 - *[Policocnemis](/source/Policocnemis)* Benjamin, 1932 - *[Triocnemis](/source/Triocnemis)* Grote, 1881 - *[Unciella](/source/Unciella)* Troubridge, 2008 - Not assigned to a subtribe - *[Allomeganephria](https://en.wikipedia.org/w/index.php?title=Allomeganephria&action=edit&redlink=1)* Ronkay, Ronkay, Gyulai & Hacker, 2010 - *[Asteroscopus](/source/Asteroscopus)* Boisduval, 1828 - *[Benedekia](https://en.wikipedia.org/w/index.php?title=Benedekia&action=edit&redlink=1)* Ronkay, Ronkay, Gyulai & Hacker, 2010 - *[Beshkovietta](https://en.wikipedia.org/w/index.php?title=Beshkovietta&action=edit&redlink=1)* Ronkay, Ronkay, Gyulai & Hacker, 2010 - *[Copitype](/source/Copitype)* Hampson, 1906 - *[Diphtherocome](/source/Diphtherocome)* Warren, 1907 - *[Fansipania](https://en.wikipedia.org/w/index.php?title=Fansipania&action=edit&redlink=1)* Ronkay, Ronkay, Gyulai & Hacker, 2010 - *[Flexivaleria](https://en.wikipedia.org/w/index.php?title=Flexivaleria&action=edit&redlink=1)* - *[Gracilisinensis](https://en.wikipedia.org/w/index.php?title=Gracilisinensis&action=edit&redlink=1)* Gyulai, Saldaitis & Vaitonis, 2017 - *[Himalaea](https://en.wikipedia.org/w/index.php?title=Himalaea&action=edit&redlink=1)* Hreblay & Ronkay, 1998 - *[Sirioba](/source/Sirioba)* Nye, 1975 - *[Speidelia](/source/Speidelia)* Ronkay, 2000 - *[Valeria](/source/Valeria_(moth))* Stephens, 1829 - *[Vigentinocome](https://en.wikipedia.org/w/index.php?title=Vigentinocome&action=edit&redlink=1)* **Not assigned to a tribe** - *[Bityla](/source/Bityla)* Walker, 1865 - *[Centrogone](/source/Centrogone)* Hampson, 1908 - *[Eutamsia](/source/Eutamsia)* Fletcher D.S., 1961 - *[Exathetis](/source/Exathetis)* Janse, 1938 - *[Heptapotamia](/source/Heptapotamia)* Alphéraky, 1882 - *[Leucosigma](/source/Leucosigma)* Druce, 1908 - *[Mazuca](/source/Mazuca)* Walker, 1866 - *[Nacna](/source/Nacna)* Fletcher, 1961 - *[Phidrimana](/source/Phidrimana)* Kononenko, 1989 - *[Thoracolopha](/source/Thoracolopha)* Turner, 1939 ## References 1. ^ [***a***](#cite_ref-Keegan2019_1-0) [***b***](#cite_ref-Keegan2019_1-1) Keegan, Kevin L.; Lafontaine, James Donald; Wahlberg, Niklas; Wagner, David L. (2019). ["Towards resolving and redefining Amphipyrinae (Lepidoptera, Noctuoidea, Noctuidae): a massively polyphyletic taxon"](https://www.biorxiv.org/content/biorxiv/early/2018/10/12/271478.full.pdf) (PDF). *Systematic Entomology*. **44** (2): 451–464. [Bibcode](/source/Bibcode_(identifier)):[2019SysEn..44..451K](https://ui.adsabs.harvard.edu/abs/2019SysEn..44..451K). [doi](/source/Doi_(identifier)):[10.1111/syen.12336](https://doi.org/10.1111%2Fsyen.12336). [S2CID](/source/S2CID_(identifier)) [92559412](https://api.semanticscholar.org/CorpusID:92559412). 1. ^ [***a***](#cite_ref-Keegan2021_2-0) [***b***](#cite_ref-Keegan2021_2-1) [***c***](#cite_ref-Keegan2021_2-2) Keegan, Kevin; Rota, Jadranka; Zahiri, Reza; Zilli, Alberto; et al. (2021). ["Toward a Stable Global Noctuidae (Lepidoptera) Taxonomy"](https://www.researchgate.net/publication/351287266). *Insect Systematics and Diversity*. **5** (3). [doi](/source/Doi_(identifier)):[10.1093/isd/ixab005](https://doi.org/10.1093%2Fisd%2Fixab005). 1. **[^](#cite_ref-itis_3-0)** ["Amphipyrinae Report"](https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=693991). *Integrated Taxonomic Information System*. Retrieved 2021-11-19. 1. **[^](#cite_ref-gbif_4-0)** ["Noctuidae"](https://www.gbif.org/species/7015). *GBIF, Global Biodiversity Information Facility*. Retrieved 2021-11-19. 1. **[^](#cite_ref-bugguide_5-0)** ["Amphipyrinae subfamily Information"](https://bugguide.net/node/view/12343). *BugGuide.net*. Retrieved 2021-11-19. ## Further reading - Grote, Aug. R.; Robinson, C. T. (1868). [*List of the Lepidoptera of North America*](https://archive.org/details/cihm_04389). American Entomological Society. [ISBN](/source/ISBN_(identifier)) [978-0-665-04389-5](https://en.wikipedia.org/wiki/Special:BookSources/978-0-665-04389-5). {{[cite book](https://en.wikipedia.org/wiki/Template:Cite_book)}}: ISBN / Date incompatibility ([help](https://en.wikipedia.org/wiki/Help:CS1_errors#invalid_isbn_date)) - Lafontaine, J. Donald; Schmidt, B. Christian (2010). ["Annotated check list of the Noctuoidea (Insecta, Lepidoptera) of North America north of Mexico"](https://doi.org/10.3897%2Fzookeys.40.414). *ZooKeys* (40): 127–47. [Bibcode](/source/Bibcode_(identifier)):[2010ZooK...40....1L](https://ui.adsabs.harvard.edu/abs/2010ZooK...40....1L). [doi](/source/Doi_(identifier)):[10.3897/zookeys.40.414](https://doi.org/10.3897%2Fzookeys.40.414). - Lafontaine, J. Donald; Schmidt, B. Christian (2015). ["Additions and corrections to the checklist of the Noctuoidea (Insecta, Lepidoptera) of North America north of Mexico, III"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4668890). *ZooKeys* (527): 227–236. [Bibcode](/source/Bibcode_(identifier)):[2015ZooK..527..127L](https://ui.adsabs.harvard.edu/abs/2015ZooK..527..127L). [doi](/source/Doi_(identifier)):[10.3897/zookeys.527.6151](https://doi.org/10.3897%2Fzookeys.527.6151). [PMC](/source/PMC_(identifier)) [4668890](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4668890). [PMID](/source/PMID_(identifier)) [26692790](https://pubmed.ncbi.nlm.nih.gov/26692790). - Pohl, Greg; Patterson, Bob; Pelham, Jonathan (2016). [Annotated taxonomic checklist of the Lepidoptera of North America, North of Mexico](https://www.researchgate.net/publication/302570819) (Report). [doi](/source/Doi_(identifier)):[10.13140/RG.2.1.2186.3287](https://doi.org/10.13140%2FRG.2.1.2186.3287). ## External links - Media related to [Amphipyrinae](https://commons.wikimedia.org/wiki/Category:Amphipyrinae) at Wikimedia Commons Taxon identifiers Amphipyrinae Wikidata: Q14552207 Wikispecies: Amphipyrinae BioLib: 53629 BugGuide: 12343 CoL: 93MRK Fauna Europaea: 449241 Fauna Europaea (new): 8444473d-58db-411a-abae-a5ce5411a90b iNaturalist: 122478 Insecta.pro: 197 ITIS: 693991 NBN: NHMSYS0000501218 NCBI: 95182 Open Tree of Life: 968470 This Amphipyrinae article is a stub. 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