# Allotheria

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{{Short description|Extinct subclass of mammals}}
{{automatic taxobox
| name = Allotheria
| fossil_range = {{fossilrange|Middle Jurassic|Eocene|earliest=Rhaetian|latest=17.5}} Possible Late Triassic record
| taxon = Allotheria
| image = Ptilodus skull.jpg
| image_caption = Skull of ''[Ptilodus](/source/Ptilodus)'' (Multituberculata)
| image2 = Adalatherium skull.svg
| image2_caption = Skull of ''[Adalatherium](/source/Adalatherium)'' (Gondwanatheria)
| authority = Marsh, 1880
| subdivision_ranks = Subgroups
| subdivision = *{{extinct}}''[Camptomus](/source/Camptomus)''
*{{extinct}}''[Fractinus](/source/Fractinus)''
*{{extinct}}?''[Cifelliodon](/source/Cifelliodon)''
*{{extinct}}''[Megaconus](/source/Megaconus)''
*{{extinct}}''[Kermackodon](/source/Kermackodon)''
*†[Gondwanatheria](/source/Gondwanatheria)
*?†[Haramiyida](/source/Haramiyida)
*†[Multituberculata](/source/Multituberculata)
}}
[[File:Xianshou linglong skull.svg|left|thumb|Skull of ''[Xianshou](/source/Xianshou),'' a [euharamiyida](/source/euharamiyida)n]]
'''Allotheria''' (from [Ancient Greek](/source/Ancient_Greek) αλλός (''allós''), meaning "other", and θηρίον (''theríon''), meaning "beast") is an [extinct](/source/extinct) clade of [theriiform](/source/Theriiformes) [mammal](/source/mammal)s known from the [Mesozoic](/source/Mesozoic) and early [Cenozoic](/source/Cenozoic). Shared characteristics of the group are the presence of lower [molariform](/source/Molar_(tooth)) [teeth](/source/Tooth_(animal)) equipped with longitudinal rows of cusps<ref>{{cite journal |url=https://www.app.pan.pl/article/item/app45-317.html |title=Review of the early allotherian mammals |first=P.M. |last= Butler |journal=Acta Palaeontologica Polonica |volume=45 |issue=4 |pages=317–342 |year=2000}}</ref> and enlarged incisors.<ref name=":0">{{Cite journal |last1=Hoffmann |first1=Simone |last2=Beck |first2=Robin M. D. |last3=Wible |first3=John R. |last4=Rougier |first4=Guillermo W. |last5=Krause |first5=David W. |date=2020-12-14 |title=Phylogenetic placement of Adalatherium hui (Mammalia, Gondwanatheria) from the Late Cretaceous of Madagascar: implications for allotherian relationships |url=https://www.tandfonline.com/doi/full/10.1080/02724634.2020.1801706 |journal=Journal of Vertebrate Paleontology |language=en |volume=40 |issue=sup1 |pages=213–234 |doi=10.1080/02724634.2020.1801706 |bibcode=2020JVPal..40S.213H |s2cid=230968231 |issn=0272-4634|url-access=subscription }}</ref> Typically, the canine teeth are also lost.<ref name=":0" /> Allotheria includes [Multituberculata](/source/Multituberculata), [Gondwanatheria](/source/Gondwanatheria) (which may be part of [Multituberculata](/source/Multituberculata), as the sister group to [Cimolodonta](/source/Cimolodonta)),<ref>{{Cite journal|last1=Krause |first1=D. W. |last2=Prasad |first2=G. V. R. |last3=von Koenigswald <!--this is correct, see paper, and used in harvnb templates, so do not change --> |first3=W. |last4=Sahni |first4=A. |last5=Grine |first5=F. E. |journal=Nature |volume=390 |issue=6659 |pages=504–507 |doi=10.1038/37343 |year=1997 |title=Cosmopolitanism among Gondwanan Late Cretaceous mammals |url=http://www.anat.stonybrook.edu/dkrause/articles/1997_lavanify.pdf |bibcode=1997Natur.390..504K |s2cid=205025618 |url-status=dead |archive-url=https://web.archive.org/web/20100611222233/http://www.anat.stonybrook.edu/dkrause/articles/1997_lavanify.pdf |archive-date=2010-06-11 }}</ref><ref name="Krause, et al">{{Cite journal|last1=Krause |first1=David W. |author-link1=David W. Krause |last2=Hoffmann |first2=Simone |last3=Wible |first3=John R. |last4=Kirk |first4=E. Christopher |last5=Schultz |first5=Julia A. |last6=von Koenigswald |first6=Wighart |last7=Groenke |first7=Joseph R. |last8=Rossie |first8=James B. |others=O'Connor, Patrick M., Seiffert, Erik R., Dumont, Elizabeth R., Holloway, Waymon L., [Rogers, Raymond R.](/source/Raymond_R._Rogers), Rahantarisoa, Lydia J., Kemp, Addison D., Andriamialison, Haingoson |date=2014-11-05 |issn=1476-4687 |title=First cranial remains of a gondwanatherian mammal reveal remarkable mosaicism |journal=Nature |publisher=Nature Publishing Group, a division of [Macmillan Publishers Limited](/source/Macmillan_Publishers_Limited) |volume=  515|issue= 7528|pages= 512–517|doi=10.1038/nature13922 |bibcode = 2014Natur.515..512K |pmid=25383528|s2cid=4395258 }}</ref><ref name="Drake">{{cite web |url=http://news.nationalgeographic.com/news/2014/11/141105-mammal-evolution-vintana-fossil-science/ |archive-url=https://web.archive.org/web/20141105183144/http://news.nationalgeographic.com/news/2014/11/141105-mammal-evolution-vintana-fossil-science/ |url-status=dead |archive-date=November 5, 2014 |title=Fossil From Dinosaur Era Reveals Big Mammal With Super Senses |date=November 5, 2014 |first=Nadia |last=Drake |author-link=Nadia Drake |access-date=November 5, 2014 |website=nationalgeographic.com |publisher=[National Geographic Society](/source/National_Geographic_Society)}}</ref><ref name="NYT-20141105-JNW">{{cite news |last=Wilford |first=John Noble |title=Fossil's Unusual Size and Location Offer Clues in Evolution of Mammals |url=https://www.nytimes.com/2014/11/06/science/madagascar-fossil-vintana-mammals-evolution.html |date=November 5, 2014 |work=[The New York Times](/source/The_New_York_Times) |access-date=November 6, 2014 }}</ref> and probably [Haramiyida](/source/Haramiyida),<ref>{{cite journal | last1 = Luo | first1 = Z.-X. | last2 = Kielan-Jaworowska | first2 = Z. | last3 = Cifelli | first3 = R.L. | year = 2002 | title = In quest for a phylogeny of Mesozoic mammals | journal = Acta Palaeontologica Polonica | volume = 47 | issue = 1 | pages = 1–78}}</ref> (sometimes only including [Euharamiyida](/source/Euharamiyida)<ref name=":0" />) although some studies have recovered haramiyidans to be basal [mammaliaform](/source/mammaliaform)s unrelated to multituberculates.<ref name="PNAS-20151116-zzxl">{{cite journal |last1=Luo |first1=Zhe-Xi |last2=Gates
|first2=Stephen M. |last3=Jenkins Jr. |first3=Farish A. |last4=Amaral |first4=William W. |last5=Shubin |first5=Neil H. |title=Mandibular and dental characteristics of Late Triassic mammaliaform Haramiyavia and their ramifications for basal mammal evolution |doi=10.1073/pnas.1519387112 |date=16 November 2015 |journal=[PNAS](/source/PNAS) |pages=E7101–E7109|bibcode=2015PNAS..112E7101L |pmc=4697399 |pmid=26630008 |volume=112|issue=51 |doi-access=free }}</ref> Allotherians are often placed as [crown group](/source/crown_group) mammals, more closely related to living [marsupial](/source/marsupial)s and [placentals](/source/placentals) ([Theria](/source/Theria)) than to [monotreme](/source/monotreme)s or [eutriconodonts](/source/Eutriconodonta),<ref name=":0" /> though some studies place the entirety of Allotheria outside of crown Mammalia.<ref>{{Cite journal |last1=Mao |first1=Fangyuan |last2=Li |first2=Zhiyu |last3=Wang |first3=Zhili |last4=Zhang |first4=Chi |last5=Rich |first5=Thomas |last6=Vickers-Rich |first6=Patricia |last7=Meng |first7=Jin |date=2024-04-03 |title=Jurassic shuotheriids show earliest dental diversification of mammaliaforms |url=https://www.nature.com/articles/s41586-024-07258-7 |journal=Nature |volume=628 |issue=8008 |pages=569–575 |language=en |doi=10.1038/s41586-024-07258-7 |pmid=38570681 |bibcode=2024Natur.628..569M |issn=0028-0836|url-access=subscription }}</ref><ref>{{Cite journal |last=Lopatin |first=A. V. |date=December 2019 |title=Modern Data on the Origin and Early Radiation of Mammals |url=http://link.springer.com/10.1134/S1062359019070082 |journal=Biology Bulletin |language=en |volume=46 |issue=7 |pages=744–750 |doi=10.1134/S1062359019070082 |issn=1062-3590|url-access=subscription }}</ref>

== References ==
{{Reflist}}

==Further reading==
Zofia Kielan-Jaworowska, Richard L. Cifelli, and Zhe-Xi Luo, ''Mammals from the Age of Dinosaurs: Origins, Evolution, and Structure'' (New York: Columbia University Press, 2004), 249.

{{Mammalia|T.|state=autocollapse}}
{{Allotheria|A.|state=autocollapse}}
{{Taxonbar|from=Q1419784}}

{{Authority control}}

Category:Allotheria
Category:Mammal taxonomy
Category:Norian first appearances
Category:Late Triassic first appearances
Category:Miocene extinctions

{{paleo-mammal-stub}}

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Adapted from the Wikipedia article [Allotheria](https://en.wikipedia.org/wiki/Allotheria) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Allotheria?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.
